FNA: Euphorbia arizonica vs. Euphorbia garberi

	Euphorbia arizonica	Euphorbia garberi
	Arizona sandmat, Arizona spurge	Garber's sandmat
Habit	Herbs, annual or short-lived perennial, with slender to slightly thickened taproot.	Herbs, perennial, with moderately thickened rootstock.
Stems	erect to ascending, 10–30 cm, uniformly pilose with glistening hairs.	ascending, sometimes slightly woody at base, not wiry, 1–3 mm diam., 15–50 cm, canescent.
Leaves	opposite; stipules distinct, deltate, (0–)0.1 mm, glabrous or with few scattered hairs; petiole 0.4–1.5 mm, pilose with glistening hairs; blade usually ovate, rarely elliptic, 3–11 × 2–7 mm, base asymmetric, one side cuneate to rounded, other side rounded to strongly cordate, margins entire, apex obtuse or acute, surfaces pilose with glistening hairs; 3-veined from base but usually only midvein conspicuous.	opposite; stipules distinct or connate at base, triangular-subulate, apex divided into 2–5 subulate to subulate-filiform segments, 0.5–0.7 mm, pilose; petiole 0.5–1.1 mm, pilose; blade ovate to oblong-elliptic, 4–9(–15) × 3–6 mm, base asymmetric, obtuse to rounded, margins usually entire, rarely very sparsely serrulate, apex usually obtuse to rounded, occasionally acute to apiculate, surfaces canescent; 3-veined from base, only midvein conspicuous.
Involucre	urceolate, 1.1–1.3 × 0.5–0.9 mm, glabrous or pilose with glistening hairs; glands 4, dark maroon, usually oblong to reniform, rarely almost circular, 0.2 × 0.2–0.4 mm; appendages white to pink, oblong, flabellate, oblong, or elliptic, 0.3–0.6 × 0.6–0.9 mm, distal margin entire or crenulate.	turbinate to campanulate, 0.6–0.9 × 0.5–0.9 mm, pilose; glands 4, brown, slightly concave, elliptic to subcircular, 0.1–0.3 × 0.2–0.4 mm; appendages white to pink, as narrow rim along gland, (0–)0.1–0.2 × 0.3–0.5 mm, surfaces pilose, distal margin entire or crenulate.
Staminate flowers	5–12.	8–20.
Pistillate flowers	ovary pilose, styles 0.5–0.6 mm, 2-fid 1/2 length.	ovary pilose; styles 0.6–0.7 mm, 2-fid 1/2 length.
Capsules	broadly ovoid to subglobose, 1.4–1.8 mm diam., pilose; columella 1.1–1.4 mm.	subglobose to broadly ovoid, 1.1–1.6 × 1.3–2.1 mm, pilose; columella 1.1–1.6 mm.
Seeds	gray to light brown, ovoid, 4-angled in cross section, 0.9–1.1 × 0.5–0.6 mm, rugose with 2–5 irregular transverse ridges that sometimes pass through abaxial keel.	gray to reddish brown, oblong-ovoid, 4-angled in cross section, 1–1.2 × 0.6–0.8 mm, flat or obscurely wrinkled.
Cyathia	solitary at distal nodes; peduncle 0.7–3.6 mm.	solitary at nodes of short, axillary branches; peduncle 0.2–0.4 mm.

	Euphorbia arizonica	Euphorbia garberi
Phenology	Flowering and fruiting year-round in response to sufficient moisture.	Flowering and fruiting year-round.
Habitat	Washes and rocky slopes, sometimes on limestone, desert scrub communities often with creosote-bush, riparian forests, mesquite woodlands, oak chaparral.	Beach dunes, coastal rock barrens, disturbed uplands and pine rocklands.
Elevation	100–1400 m. (300–4600 ft.)	0–10 m. (0–0 ft.)
Distribution	AZ; CA; NM; TX; Mexico (Baja California, Baja California Sur, Chihuahua, Coahuila, Durango, Sonora) [WildflowerSearch map] [BONAP county map]	FL [BONAP county map]
Discussion	Euphorbia arizonica is distinctive and easily recognized by its glistening, translucent hairs that appear somewhat glutinous and are most apparent on the stems. (Discussion copyrighted by Flora of North America; reprinted with permission.)	Euphorbia garberi appears to be an interspecific hybrid, but its parentage is not entirely clear (Y. Yang and P. E. Berry 2011). The taxon is closely related to E. blodgettii, E. porteriana, and E. serpens. It is restricted to Collier, Miami-Dade, and Monroe counties in southern Florida and is federally listed as threatened due to impacts from habitat loss, fire suppression, and invasive species. A. Herndon (1993) synonymized E. porteriana var. keyensis under E. garberi because of its uniformly pilose capsules, and that treatment is followed here. (Discussion copyrighted by Flora of North America; reprinted with permission.)
Source	FNA vol. 12, p. 260.	FNA vol. 12, p. 268.
Parent taxa	Euphorbiaceae > Euphorbia > sect. Anisophyllum	Euphorbiaceae > Euphorbia > sect. Anisophyllum
Sibling taxa	E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana	E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana
Synonyms	Chamaesyce arizonica	Chamaesyce adicioides, C. brachypoda, C. garberi, C. keyensis, C. mosieri, C. porteriana var. keyensis, E. porteriana var. keyensis
Name authority	Engelmann: in W. H. Emory, Rep. U.S. Mex. Bound. 2(1): 186. (1859)	Engelmann ex Chapman: Fl. South. U.S. ed. 2, 646. (1883)
Web links	Local floras: CA Local Web sites: CalFlora, CalPhotos WildflowerSearch iNaturalist (observations) USDA Plants Database LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search	WildflowerSearch iNaturalist (observations) USDA Plants Database LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search

Euphorbia garberi

Arizona sandmat, Arizona spurge

Garber's sandmat

Habit

Herbs, annual or short-lived perennial, with slender to slightly thickened taproot.

Herbs, perennial, with moderately thickened rootstock.

Stems

erect to ascending, 10–30 cm, uniformly pilose with glistening hairs.

ascending, sometimes slightly woody at base, not wiry, 1–3 mm diam., 15–50 cm, canescent.

Leaves

opposite;

stipules distinct, deltate, (0–)0.1 mm, glabrous or with few scattered hairs;

petiole 0.4–1.5 mm, pilose with glistening hairs;

blade usually ovate, rarely elliptic, 3–11 × 2–7 mm, base asymmetric, one side cuneate to rounded, other side rounded to strongly cordate, margins entire, apex obtuse or acute, surfaces pilose with glistening hairs; 3-veined from base but usually only midvein conspicuous.

opposite;

stipules distinct or connate at base, triangular-subulate, apex divided into 2–5 subulate to subulate-filiform segments, 0.5–0.7 mm, pilose;

petiole 0.5–1.1 mm, pilose;

blade ovate to oblong-elliptic, 4–9(–15) × 3–6 mm, base asymmetric, obtuse to rounded, margins usually entire, rarely very sparsely serrulate, apex usually obtuse to rounded, occasionally acute to apiculate, surfaces canescent; 3-veined from base, only midvein conspicuous.

Involucre

urceolate, 1.1–1.3 × 0.5–0.9 mm, glabrous or pilose with glistening hairs;

glands 4, dark maroon, usually oblong to reniform, rarely almost circular, 0.2 × 0.2–0.4 mm;

appendages white to pink, oblong, flabellate, oblong, or elliptic, 0.3–0.6 × 0.6–0.9 mm, distal margin entire or crenulate.

turbinate to campanulate, 0.6–0.9 × 0.5–0.9 mm, pilose;

glands 4, brown, slightly concave, elliptic to subcircular, 0.1–0.3 × 0.2–0.4 mm;

appendages white to pink, as narrow rim along gland, (0–)0.1–0.2 × 0.3–0.5 mm, surfaces pilose, distal margin entire or crenulate.

Staminate flowers

5–12.

8–20.

Pistillate flowers

ovary pilose, styles 0.5–0.6 mm, 2-fid 1/2 length.

ovary pilose;

styles 0.6–0.7 mm, 2-fid 1/2 length.

Capsules

broadly ovoid to subglobose, 1.4–1.8 mm diam., pilose;

columella 1.1–1.4 mm.

subglobose to broadly ovoid, 1.1–1.6 × 1.3–2.1 mm, pilose;

columella 1.1–1.6 mm.

Seeds

gray to light brown, ovoid, 4-angled in cross section, 0.9–1.1 × 0.5–0.6 mm, rugose with 2–5 irregular transverse ridges that sometimes pass through abaxial keel.

gray to reddish brown, oblong-ovoid, 4-angled in cross section, 1–1.2 × 0.6–0.8 mm, flat or obscurely wrinkled.

Cyathia

solitary at distal nodes;

peduncle 0.7–3.6 mm.

solitary at nodes of short, axillary branches;

peduncle 0.2–0.4 mm.

Euphorbia arizonica

Euphorbia garberi

Phenology

Flowering and fruiting year-round in response to sufficient moisture.

Flowering and fruiting year-round.

Habitat

Washes and rocky slopes, sometimes on limestone, desert scrub communities often with creosote-bush, riparian forests, mesquite woodlands, oak chaparral.

Beach dunes, coastal rock barrens, disturbed uplands and pine rocklands.

Elevation

100–1400 m. (300–4600 ft.)

0–10 m. (0–0 ft.)

Distribution

AZ; CA; NM; TX; Mexico (Baja California, Baja California Sur, Chihuahua, Coahuila, Durango, Sonora)

[WildflowerSearch map]

[BONAP county map]

Discussion

Euphorbia arizonica is distinctive and easily recognized by its glistening, translucent hairs that appear somewhat glutinous and are most apparent on the stems.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Euphorbia garberi appears to be an interspecific hybrid, but its parentage is not entirely clear (Y. Yang and P. E. Berry 2011). The taxon is closely related to E. blodgettii, E. porteriana, and E. serpens. It is restricted to Collier, Miami-Dade, and Monroe counties in southern Florida and is federally listed as threatened due to impacts from habitat loss, fire suppression, and invasive species. A. Herndon (1993) synonymized E. porteriana var. keyensis under E. garberi because of its uniformly pilose capsules, and that treatment is followed here.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source

FNA vol. 12, p. 260.

FNA vol. 12, p. 268.

Parent taxa

Euphorbiaceae > Euphorbia > sect. Anisophyllum

Sibling taxa

E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana

E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana

Synonyms

Chamaesyce arizonica

Chamaesyce adicioides, C. brachypoda, C. garberi, C. keyensis, C. mosieri, C. porteriana var. keyensis, E. porteriana var. keyensis

Name authority

Engelmann: in W. H. Emory, Rep. U.S. Mex. Bound. 2(1): 186. (1859)

Engelmann ex Chapman: Fl. South. U.S. ed. 2, 646. (1883)

Web links

Unless otherwise noted, all data comes from the online version of the Flora of North America, who have made most of their content available via a Creative Commons license (CC BY 4.0).

Choose a data source: Flora of North America

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Euphorbia arizonica

Euphorbia garberi

Euphorbia arizonica

Euphorbia garberi