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Arizona sandmat, Arizona spurge

fork spurge

Habit Herbs, annual or short-lived perennial, with slender to slightly thickened taproot. Herbs, annual, with slender, fibrous taproot.
Stems

erect to ascending, 10–30 cm, uniformly pilose with glistening hairs.

erect, 20–70 cm, glabrous or with few scattered spreading hairs;

branches arcuate, branching appearing dichotomous.

Leaves

opposite;

stipules distinct, deltate, (0–)0.1 mm, glabrous or with few scattered hairs;

petiole 0.4–1.5 mm, pilose with glistening hairs;

blade usually ovate, rarely elliptic, 3–11 × 2–7 mm, base asymmetric, one side cuneate to rounded, other side rounded to strongly cordate, margins entire, apex obtuse or acute, surfaces pilose with glistening hairs; 3-veined from base but usually only midvein conspicuous.

usually alternate, occasionally opposite at proximalmost node;

petiole 15–49 mm, glabrous;

blade usually ovate, rarely oblong or elliptic, 13–54 × 7–38 mm, base usually rounded to broadly cuneate, rarely truncate, margins finely serrulate, apex obtuse, surfaces glabrous or with few scattered hairs;

venation pinnate, midvein prominent.

Involucre

urceolate, 1.1–1.3 × 0.5–0.9 mm, glabrous or pilose with glistening hairs;

glands 4, dark maroon, usually oblong to reniform, rarely almost circular, 0.2 × 0.2–0.4 mm;

appendages white to pink, oblong, flabellate, oblong, or elliptic, 0.3–0.6 × 0.6–0.9 mm, distal margin entire or crenulate.

tubular or obconic, 1–1.7 × 0.7–1.4 mm, glabrous except for few hairs on lobes;

involucral lobes divided into several linear, smooth lobes;

glands 1(–3), greenish, sessile and broadly attached, 0.3–0.4 × 0.4–0.8 mm, opening oblong to subcircular, glabrous;

appendages petaloid, white, elliptic, oblong, transversely oblong, or forming thin, lunate rim on gland margin, not incurved and covering glands, 0.3–0.9 × 0.6–1.3 mm, entire, undulate or slightly lobed, glabrous.

Staminate flowers

5–12.

20–30.

Pistillate flowers

ovary pilose, styles 0.5–0.6 mm, 2-fid 1/2 length.

ovary glabrous;

styles 0.6–1 mm, 2-fid 1/2 length.

Capsules

broadly ovoid to subglobose, 1.4–1.8 mm diam., pilose;

columella 1.1–1.4 mm.

oblate, 2.8–3.1 × 3.6–4.5 mm, glabrous;

columella 1.9–2.4 mm.

Seeds

gray to light brown, ovoid, 4-angled in cross section, 0.9–1.1 × 0.5–0.6 mm, rugose with 2–5 irregular transverse ridges that sometimes pass through abaxial keel.

brown to blackish, ovoid, rounded in cross section, 1.9–2.4 × 1.5–1.8 mm, irregularly and coarsely tuberculate;

caruncle absent or rudimentary.

Cyathia

solitary at distal nodes;

peduncle 0.7–3.6 mm.

peduncle 0.9–3.5(–6.2) mm.

Cyathial

arrangement: terminal dichasial branches 2, few-branched (weakly defined);

pleiochasial bracts 2–3, opposite or whorled, wholly green, similar in shape and size to distal leaves;

dichasial bracts smaller than distal leaves, often white at base.

Euphorbia arizonica

Euphorbia bifurcata

Phenology Flowering and fruiting year-round in response to sufficient moisture. Flowering and fruiting summer–fall.
Habitat Washes and rocky slopes, sometimes on limestone, desert scrub communities often with creosote-bush, riparian forests, mesquite woodlands, oak chaparral. Riparian areas with cottonwoods and willows, pinyon pine woodlands, pine-oak forests, Douglas fir forests with pines.
Elevation 100–1400 m. (300–4600 ft.) 1900–2300 m. (6200–7500 ft.)
Distribution
from FNA
AZ; CA; NM; TX; Mexico (Baja California, Baja California Sur, Chihuahua, Coahuila, Durango, Sonora)
[WildflowerSearch map]
[BONAP county map]
from FNA
NM; TX; Mexico; Central America (Guatemala)
[BONAP county map]
Discussion

Euphorbia arizonica is distinctive and easily recognized by its glistening, translucent hairs that appear somewhat glutinous and are most apparent on the stems.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Euphorbia bifurcata is found in the mountains of southern New Mexico (Doña Ana, Grant, Lincoln, Otero, and Sierra counties) and trans-Pecos Texas (Brewster, Jeff Davis, and Presidio counties).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 12, p. 260. FNA vol. 12, p. 319.
Parent taxa Euphorbiaceae > Euphorbia > sect. Anisophyllum Euphorbiaceae > Euphorbia > sect. Poinsettia
Sibling taxa
E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana
E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana
Synonyms Chamaesyce arizonica
Name authority Engelmann: in W. H. Emory, Rep. U.S. Mex. Bound. 2(1): 186. (1859) Engelmann: in W. H. Emory, Rep. U.S. Mex. Bound. 2(1): 190. (1859)
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