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candelilla, wax plant

Wright's spurge

Habit Shrubs, with much-branched, fleshy rootstock. Herbs, perennial with cylindric rootstock or elongated tubers.
Stems

erect, few branched, 25–50(–100) cm, glabrous or puberulent, pencil-like, in age covered with flaky, exfoliating layer of wax.

erect, branched, densely clumped, previous year’s dead stems often persistent, 20–50 cm, glabrous.

Leaves

alternate usually caducous, sometimes persisting;

stipules 0.4–0.5 mm;

petiole absent;

blade ovate to deltate-subulate, 2.5–4 × 1 mm, thick, fleshy, base usually rounded and swollen, rarely cuneate, margins entire, apex acute, surfaces puberulent, adaxial sometimes canescent;

venation inconspicuous.

alternate, persisting, spreading or ascending;

stipules 0.1–0.3 mm;

petiole absent;

blade linear to linear-filiform, (17–)20–40 × 1–2.5 mm, base cuneate, slightly sheathing stem, margins entire, apex broadly acute to rounded, abaxial surface glabrous or sparsely villous, adaxial surface glabrous;

venation obscure, only midvein conspicuous on wider leaves.

Involucre

campanulate, 1.6–2.2 × 1.6–1.9 mm, puberulent to canescent;

glands 5, pinkish, narrowly oblong to reniform, 0.3–0.4 × 0.8–1 mm;

appendages white to pink, ovate, oblong, or transversely oblong, 1.3–2.5 × 1.4–2.5 mm, usually erose, rarely entire.

campanulate, 1.5–2 × 1.8–2.5 mm, pilose;

glands 5, green, broadly elliptic, 0.7–0.8 × 1 mm;

appendages white to pink, orbiculate, 0.5–1 × 1.3–1.8 mm, coarsely erose.

Staminate flowers

50–70.

20–25.

Pistillate flowers

ovary glabrous;

styles 0.9–1.1 mm, 2-fid nearly entire length.

ovary strigose to tomentose;

styles 1 mm, 2-fid at apex.

Capsules

oblong to ovoid, 3.9–4.2 × 3.6–3.9 mm, glabrous;

columella 3.1–3.3 mm.

depressed-globose, 1 locule usually aborting, 2.5(–3) × 2.7–3.3(–5) mm, sparsely tomentose;

columella 2.6–3 mm.

Seeds

whitish gray, narrowly ovoid, 2.4–3.1 × 1.4–1.6 mm, irregularly rugose-tuberculate;

caruncle crescent-shaped, 0.3–0.6 × 0.6–0.8 mm.

gray-green to gray-brown, globose-ovoid, 2.2–2.9 × 1.8–2 mm, shallowly and obscurely pitted;

caruncle absent.

Cyathia

in axillary congested cymes, near branch tips or solitary at distal nodes;

peduncle 0–1 mm, lanulose.

usually in terminal dichasia, rarely pleiochasia;

peduncle (3–)5–15 mm, all peduncles and cyathia persistent on plant, glabrous.

Euphorbia antisyphilitica

Euphorbia wrightii

Phenology Flowering and fruiting year-round in response to sufficient rainfall. Flowering and fruiting spring–fall.
Habitat Desert scrub, frequently on limestone substrates. Open grasslands and uplands, often on limestone outcrops.
Elevation 100–1200 m. (300–3900 ft.) 500–1000 m. (1600–3300 ft.)
Distribution
from FNA
NM; TX; Mexico
[WildflowerSearch map]
[BONAP county map]
from FNA
TX; Mexico (Coahuila)
[BONAP county map]
Discussion

Euphorbia antisyphilitica is the only pencil-stemmed species of Euphorbia occurring in the flora area. The species is characteristic of the Chihuahuan Desert scrub of Mexico from Chihuahua and Coahuila south to Hidalgo and Querétaro, and barely enters into the United States in southern New Mexico (Doña Ana and Lincoln counties) and southwest (Brewster, Hudspeth, Presidio, and Terrell counties) and south (Starr and Webb counties) Texas. The stems are covered in a conspicuous coat of exfoliating wax, and the plants historically have been harvested for this product, although the practice is much less prevalent now. The specific epithet refers to its traditional medicinal use in treating sexually transmitted infections.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Within the flora area, Euphorbia wrightii is endemic to the western Edwards Plateau and adjacent rolling plains in western Texas.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 12, p. 243. FNA vol. 12, p. 251.
Parent taxa Euphorbiaceae > Euphorbia > sect. Alectoroctonum Euphorbiaceae > Euphorbia > sect. Alectoroctonum
Sibling taxa
E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana
E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. yaquiana
Name authority Zuccarini: Flora 15(2, Beibl.): 58. (1832) Torrey & A. Gray: in War Department [U.S.], Pacif. Railr. Rep. 2(4): 174. (1857)
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