Euphorbia antisyphilitica |
Euphorbia indivisa |
|
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candelilla, wax plant |
royal sandmat |
|
Habit | Shrubs, with much-branched, fleshy rootstock. | Herbs, annual or short-lived perennial, with slender taproot to thickened and woody rootstock. |
Stems | erect, few branched, 25–50(–100) cm, glabrous or puberulent, pencil-like, in age covered with flaky, exfoliating layer of wax. |
prostrate, usually mat-forming, terete to slightly flattened, 40 cm, lower surface glabrous, upper surface strigillose, pilose or villous. |
Leaves | alternate usually caducous, sometimes persisting; stipules 0.4–0.5 mm; petiole absent; blade ovate to deltate-subulate, 2.5–4 × 1 mm, thick, fleshy, base usually rounded and swollen, rarely cuneate, margins entire, apex acute, surfaces puberulent, adaxial sometimes canescent; venation inconspicuous. |
opposite; stipules distinct, entire or divided into 3–4 subulate to filiform segments, 0.8–2 mm, usually pilose, rarely glabrous; petiole 0.5–1 mm, pilose to villous; blade oblong, ovate or narrowly obovate, 3–10(–12) × 2–6 mm, base strongly asymmetric, hemicordate, margins serrulate, apex obtuse to subacute, surfaces glabrous or slightly pilose; 3-veined from base, often only midvein conspicuous. |
Involucre | campanulate, 1.6–2.2 × 1.6–1.9 mm, puberulent to canescent; glands 5, pinkish, narrowly oblong to reniform, 0.3–0.4 × 0.8–1 mm; appendages white to pink, ovate, oblong, or transversely oblong, 1.3–2.5 × 1.4–2.5 mm, usually erose, rarely entire. |
narrowly turbinate, 1–1.2 × 0.4–0.7 mm, pilose; glands 4, yellow to pink, unequal, proximal pair oblong or linear, 0.1 × 0.3–0.4(–0.6) mm, distal pair oblong or subcircular, 0.1 × 0.1–0.2 mm; appendages pink to reddish, unequal, on proximal glands oblique, 0.4–0.8(–1) × 0.8–1.4(–2) mm, on distal glands symmetric, 0.2–0.5 × 0.2–0.5 mm, slightly undulate to slightly crenate. |
Staminate flowers | 50–70. |
5–15. |
Pistillate flowers | ovary glabrous; styles 0.9–1.1 mm, 2-fid nearly entire length. |
ovary pilose to strigillose in parts, glabrous in other parts; styles 0.8–1.3 mm, usually unbranched, rarely 2-fid at apex, filiform. |
Capsules | oblong to ovoid, 3.9–4.2 × 3.6–3.9 mm, glabrous; columella 3.1–3.3 mm. |
ovoid-triangular, 1.2–1.5 × 1–1.4 mm, pilose to strigillose in parts, glabrous in other parts; columella 1–1.3 mm. |
Seeds | whitish gray, narrowly ovoid, 2.4–3.1 × 1.4–1.6 mm, irregularly rugose-tuberculate; caruncle crescent-shaped, 0.3–0.6 × 0.6–0.8 mm. |
brown to light gray, ovoid, 4-angled in cross section, 0.8–1 × 0.4–0.5 mm, with 4 or 5 deep transverse sulci alternating with low transverse ridges. |
Cyathia | in axillary congested cymes, near branch tips or solitary at distal nodes; peduncle 0–1 mm, lanulose. |
usually in small cymose clusters on congested, axillary branches; peduncle rudimentary or to 0.2 mm. |
Euphorbia antisyphilitica |
Euphorbia indivisa |
|
Phenology | Flowering and fruiting year-round in response to sufficient rainfall. | Flowering and fruiting summer–fall. |
Habitat | Desert scrub, frequently on limestone substrates. | Grasslands, oak forests, oak-mesquite woodlands, oak-juniper communities, rarely entering desert scrub. |
Elevation | 100–1200 m. (300–3900 ft.) | 1000–2000 m. (3300–6600 ft.) |
Distribution |
NM; TX; Mexico
|
AZ; NM; TX; Mexico
|
Discussion | Euphorbia antisyphilitica is the only pencil-stemmed species of Euphorbia occurring in the flora area. The species is characteristic of the Chihuahuan Desert scrub of Mexico from Chihuahua and Coahuila south to Hidalgo and Querétaro, and barely enters into the United States in southern New Mexico (Doña Ana and Lincoln counties) and southwest (Brewster, Hudspeth, Presidio, and Terrell counties) and south (Starr and Webb counties) Texas. The stems are covered in a conspicuous coat of exfoliating wax, and the plants historically have been harvested for this product, although the practice is much less prevalent now. The specific epithet refers to its traditional medicinal use in treating sexually transmitted infections. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Euphorbia indivisa is characteristic of grasslands and oak woodlands from extreme western Texas to southeastern Arizona. The species is often treated as a synonym of E. dioeca Kunth, but the two species are readily separable on the basis of their seeds. The seeds of E. indivisa possess deep transverse sulci, whereas those of E. dioeca are merely rippled or with low transverse ridges. Euphorbia dioeca is a weedy species that occurs widely throughout tropical America but has yet to be encountered within the flora area. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 12, p. 243. | FNA vol. 12, p. 273. |
Parent taxa | Euphorbiaceae > Euphorbia > sect. Alectoroctonum | Euphorbiaceae > Euphorbia > sect. Anisophyllum |
Sibling taxa | ||
Synonyms | E. dioeca var. indivisa, Chamaesyce indivisa | |
Name authority | Zuccarini: Flora 15(2, Beibl.): 58. (1832) | (Engelmann) Tidestrom: Proc. Biol. Soc. Wash. 48: 40. (1935) |
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