Euphorbia alta |
Euphorbia stictospora |
|
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giant spurge, roughpod spurge, tall spurge, warty spurge |
mat spurge, narrow-seed spurge, slimseed sandmat |
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Habit | Herbs, annual or biennial, with taproot. | Herbs, annual, with taproot. |
Stems | erect, branched, 20–60 cm, glabrous. |
prostrate, often mat-forming, occasionally with ascending tips, 5–45 cm, densely and evenly pilose to lanate. |
Leaves | petiole 0–1 mm; blade oblong-spatulate, 20–50 × 7–18 mm, base broadly attenuate, margins serrulate, apex rounded to obtuse, surfaces glabrous, ± glaucous; venation pinnate, midvein prominent. |
opposite; stipules distinct or connate basally on one side of stem, entire or irregularly toothed or fringed, 0.5–1.2 mm, pilose to lanate; petiole 0.3–1.5 mm, pilose to lanate; blade usually oblong to oblong-obovate, occasionally nearly circular, 3–10(–15) × 2–5(–10) mm, base asymmetric, one side usually angled or rounded and other truncate-auriculate, margins minutely or conspicuously serrulate at least toward apex, apex usually broadly rounded to broadly acute, occasionally emarginate, abaxial surface often ± lighter green and without reddish spot, both surfaces sparsely to moderately pilose to lanate; 3-veined from base or venation obscure. |
Involucre | narrowly campanulate, 0.8–1.1 × 1.1–1.3 mm, glabrous; glands 4, elliptic, 0.3–0.5 × 0.5–0.7 mm; horns absent. |
obconic, 0.7–1 × 0.4–0.6 mm, moderately to densely pilose to lanate; glands 4, reddish, ± unequal, oblong, 0.1 × 0.1–0.3 mm; appendages white to strongly pinkish or reddish tinged, often unequal, sometimes 1 to all absent, 0–0.3 × 0–0.4 mm, 3-lobed or rudimentarily 1-lobed, distal margin crenate. |
Staminate flowers | 5–10. |
3–9. |
Pistillate flowers | ovary glabrous; styles 0.5–0.9 mm, 2-fid. |
ovary pilose to villous, hairs occasionally slightly appressed; styles 0.2–0.5 mm, unbranched or inconspicuously notched at tip. |
Capsules | depressed-globose, 2–3 × 2.5–3.5 mm, 3-lobed; cocci rounded, papillate, papillae 0.2–0.5 mm, glabrous; columella 1.5–1.9 mm. |
ovoid, 1.6–2.3 × 1.4–1.5 mm, moderately to densely villous with hairs usually slightly appressed, pubescence often concentrated on proximal 1/2 or along lobes; columella 1.5–2 mm. |
Seeds | purple-black, ovoid, 1.6–2 × 1.3–1.7 mm, reticulate and areolate; caruncle reniform, flat, 0.5 × 0.7 mm. |
light to dark brown, usually mottled, sometimes with thin, white coating, often wearing away irregularly, narrowly oblong-ovoid to ellipsoid, 3–4-angled in cross section, 1–1.5 × 0.5–0.6 mm, with short, irregularly interrupted furrows, appearing partially and irregularly few-ridged. |
Cyathial | arrangement: terminal pleiochasial branches 3, 2–3 times 2-branched; pleiochasial bracts elliptic-oblanceolate to oblong, similar in size to distal leaves; dichasial bracts distinct, broadly ovate to orbiculate/reniform, base obtuse, margins serrulate, apex obtuse to rounded and often mucronulate; axillary cymose branches 6–20(–25). |
|
Cyathia | peduncle 0.5–1 mm. |
solitary at leaf nodes or in small, cymose clusters on congested, axillary branches; peduncle 0.7–2.5 mm. |
Euphorbia alta |
Euphorbia stictospora |
|
Phenology | Flowering and fruiting summer. | Flowering and fruiting midsummer–early fall. |
Habitat | Montane pine-oak and mixed conifer forests, disturbed roadsides, logged areas. | Open disturbed areas, rocky slopes. |
Elevation | 1500–3000 m. (4900–9800 ft.) | 100–2100 m. (300–6900 ft.) |
Distribution |
AZ; NM; Mexico
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AZ; CO; IA; KS; MO; ND; NE; NM; OK; SD; TX; WY; Mexico (Chihuahua, Durango, San Luis Potosí)
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Discussion | Euphorbia alta is a montane species from southern Arizona, New Mexico, and northern and central Mexico that is very similar to and sometimes difficult to distinguish from E. spathulata. Euphorbia alta tends to be a robust biennial, whereas E. spathulata is strictly annual. The most consistent characteristic to separate these two species is that the ovaries and capsules of E. alta are distinctly papillate, with the papillae rising sharply above the surface, whereas the ovaries and capsules of E. spathulata are merely verrucose, with the protuberances lower and rounded. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Euphorbia stictospora has been recorded from New York, but this disjunct occurrence likely represents a waif or misidentification. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 12, p. 298. | FNA vol. 12, p. 289. |
Parent taxa | Euphorbiaceae > Euphorbia > subg. Esula | Euphorbiaceae > Euphorbia > sect. Anisophyllum |
Sibling taxa | ||
Synonyms | Tithymalus altus | Chamaesyce stictospora |
Name authority | Norton: N. Amer. Euphorbia, 24, plate 24. (1899) | Engelmann: in W. H. Emory, Rep. U.S. Mex. Bound. 2(1): 187. (1859) |
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