Euphorbia alta |
Euphorbia porteriana |
|
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giant spurge, roughpod spurge, tall spurge, warty spurge |
Porter's sandmat |
|
Habit | Herbs, annual or biennial, with taproot. | Herbs, usually annual, sometimes perennial, with slender to occasionally thickened taproot, 5 mm diam.. |
Stems | erect, branched, 20–60 cm, glabrous. |
erect to ascending, sometimes slightly woody at base, not mat-forming, 15–60 cm, usually glabrous, young branches rarely very sparsely short pubescent. |
Leaves | petiole 0–1 mm; blade oblong-spatulate, 20–50 × 7–18 mm, base broadly attenuate, margins serrulate, apex rounded to obtuse, surfaces glabrous, ± glaucous; venation pinnate, midvein prominent. |
opposite; stipules usually distinct, occasionally connate basally, triangular, entire or 2–4 parted, apically ciliate, 0.3–0.4 mm, usually glabrous, rarely very sparsely short pubescent; petiole 0.5–1.5 mm, usually glabrous, rarely very sparsely short pubescent; blade ovate, oval, elliptic, oblong-elliptic, or linear-elliptic, 4–12 × 1–7 mm, base asymmetric, rounded or subcordate, margins usually entire, rarely sparsely serrate, apex obtuse to acute, sometimes mucronate, surfaces abaxially often purple or red, adaxially green, usually glabrous, rarely very sparsely short pubescent; 3-veined from base, only midvein conspicuous. |
Involucre | narrowly campanulate, 0.8–1.1 × 1.1–1.3 mm, glabrous; glands 4, elliptic, 0.3–0.5 × 0.5–0.7 mm; horns absent. |
turbinate to campanulate, 0.7–1 × 0.8–1 mm, usually glabrous, rarely very sparsely short pubescent; glands 4, brown, slightly concave, elliptic-oblong, 0.1–0.2 × 0.3–0.6 mm; appendages white or pink to dark red, usually oblong or flabellate, occasionally rudimentary and forming narrow rim at edge of gland, (0–)0.1–0.2 × 0.4–1 mm, distal margin entire or crenulate. |
Staminate flowers | 5–10. |
8–14. |
Pistillate flowers | ovary glabrous; styles 0.5–0.9 mm, 2-fid. |
ovary glabrous, subtended by triangular calyxlike structure; styles 0.4–0.6 mm, 2-fid 1/2 length. |
Capsules | depressed-globose, 2–3 × 2.5–3.5 mm, 3-lobed; cocci rounded, papillate, papillae 0.2–0.5 mm, glabrous; columella 1.5–1.9 mm. |
broadly ovoid, 1–1.5 × 1.7–2.1 mm, glabrous; columella 1–1.4 mm. |
Seeds | purple-black, ovoid, 1.6–2 × 1.3–1.7 mm, reticulate and areolate; caruncle reniform, flat, 0.5 × 0.7 mm. |
reddish brown, ovoid, 4-angled in cross section, 0.7–1 × 0.6–0.7 mm, smooth or obscurely wrinkled. |
Cyathial | arrangement: terminal pleiochasial branches 3, 2–3 times 2-branched; pleiochasial bracts elliptic-oblanceolate to oblong, similar in size to distal leaves; dichasial bracts distinct, broadly ovate to orbiculate/reniform, base obtuse, margins serrulate, apex obtuse to rounded and often mucronulate; axillary cymose branches 6–20(–25). |
|
Cyathia | peduncle 0.5–1 mm. |
solitary at distal nodes; peduncle 0.5–1.1 mm. |
Euphorbia alta |
Euphorbia porteriana |
|
Phenology | Flowering and fruiting summer. | Flowering and fruiting year-round. |
Habitat | Montane pine-oak and mixed conifer forests, disturbed roadsides, logged areas. | Pine rocklands, coastal scrub, open hammocks. |
Elevation | 1500–3000 m. (4900–9800 ft.) | 0–10 m. (0–0 ft.) |
Distribution |
AZ; NM; Mexico
|
FL |
Discussion | Euphorbia alta is a montane species from southern Arizona, New Mexico, and northern and central Mexico that is very similar to and sometimes difficult to distinguish from E. spathulata. Euphorbia alta tends to be a robust biennial, whereas E. spathulata is strictly annual. The most consistent characteristic to separate these two species is that the ovaries and capsules of E. alta are distinctly papillate, with the papillae rising sharply above the surface, whereas the ovaries and capsules of E. spathulata are merely verrucose, with the protuberances lower and rounded. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Euphorbia porteriana is found in Miami-Dade County and on Big Pine Key, Monroe County. This species is closely related to E. blodgettii, E. garberi, and E. serpens. The capsules of E. porteriana are almost always completely glabrous, but the authors have seen three specimens with just a few scattered hairs on the keels. A. Herndon (1993) synonymized E. porteriana var. keyensis under E. garberi because of its uniformly hairy capsules, and that treatment is followed here. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 12, p. 298. | FNA vol. 12, p. 285. |
Parent taxa | Euphorbiaceae > Euphorbia > subg. Esula | Euphorbiaceae > Euphorbia > sect. Anisophyllum |
Sibling taxa | ||
Synonyms | Tithymalus altus | Chamaesyce porteriana, C. porteriana var. scoparia, C. scoparia |
Name authority | Norton: N. Amer. Euphorbia, 24, plate 24. (1899) | (Small) Oudejans: Phytologia 67: 48. (1989) |
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