Euphorbia alta |
Euphorbia ipecacuanhae |
|
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giant spurge, roughpod spurge, tall spurge, warty spurge |
American ipecac, Carolina ipecac, ipecac spurge |
|
Habit | Herbs, annual or biennial, with taproot. | Herbs, perennial, with deep, stout rootstock. |
Stems | erect, branched, 20–60 cm, glabrous. |
decumbent or slightly ascending, branched, often densely clumped, previous year's dead stems not persistent, 17–27 cm, usually glabrous, rarely sparsely villous. |
Leaves | petiole 0–1 mm; blade oblong-spatulate, 20–50 × 7–18 mm, base broadly attenuate, margins serrulate, apex rounded to obtuse, surfaces glabrous, ± glaucous; venation pinnate, midvein prominent. |
alternate; stipules 0.1–0.2 mm; petiole (0–)1–2 mm, glabrous; blade usually linear, obovate, or oblanceolate to orbiculate, rarely filiform, proximal greatly reduced, scalelike, 15–70 × 1.5–13 mm gradually smaller proximally, base cuneate, margins entire, apex rounded, broadly acute, or emarginate, surfaces glabrous; venation occasionally obscure on smaller leaves, midvein conspicuous. |
Involucre | narrowly campanulate, 0.8–1.1 × 1.1–1.3 mm, glabrous; glands 4, elliptic, 0.3–0.5 × 0.5–0.7 mm; horns absent. |
yellow or yellow-green, hemispheric, 1–1.2 × 2–2.4 mm, glabrous; glands 5, yellow or yellow-green, obovate or elliptic, 0.7–0.8 × 1–1.2 mm; appendages white or green, often forming narrow rim around distal margin of gland, 0–0.2 mm, entire. |
Staminate flowers | 5–10. |
10–20. |
Pistillate flowers | ovary glabrous; styles 0.5–0.9 mm, 2-fid. |
ovary glabrous; styles 0.4–0.8 mm, 2-fid at apex. |
Capsules | depressed-globose, 2–3 × 2.5–3.5 mm, 3-lobed; cocci rounded, papillate, papillae 0.2–0.5 mm, glabrous; columella 1.5–1.9 mm. |
globose, 2.3–3.4 × 3.5–4.2 mm, glabrous; columella 3–3.1 mm. |
Seeds | purple-black, ovoid, 1.6–2 × 1.3–1.7 mm, reticulate and areolate; caruncle reniform, flat, 0.5 × 0.7 mm. |
white or brown, ovoid, 2.3–2.5 × 1.5 mm, angular, with 5 longitudinal ridges, shallowly and irregularly pitted; caruncle absent. |
Cyathial | arrangement: terminal pleiochasial branches 3, 2–3 times 2-branched; pleiochasial bracts elliptic-oblanceolate to oblong, similar in size to distal leaves; dichasial bracts distinct, broadly ovate to orbiculate/reniform, base obtuse, margins serrulate, apex obtuse to rounded and often mucronulate; axillary cymose branches 6–20(–25). |
|
Cyathia | peduncle 0.5–1 mm. |
usually in terminal dichasia, sometimes pleiochasia; peduncle 10–50(–70) mm, glabrous. |
Euphorbia alta |
Euphorbia ipecacuanhae |
|
Phenology | Flowering and fruiting summer. | Flowering and fruiting early spring–early summer. |
Habitat | Montane pine-oak and mixed conifer forests, disturbed roadsides, logged areas. | Pine and pine-oak savannas, pine-oak sand hills, turkey oak scrub, open sand habitats. |
Elevation | 1500–3000 m. (4900–9800 ft.) | 0–150 m. (0–500 ft.) |
Distribution |
AZ; NM; Mexico
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CT; DC; DE; GA; MD; NC; NJ; NY; PA; SC; VA
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Discussion | Euphorbia alta is a montane species from southern Arizona, New Mexico, and northern and central Mexico that is very similar to and sometimes difficult to distinguish from E. spathulata. Euphorbia alta tends to be a robust biennial, whereas E. spathulata is strictly annual. The most consistent characteristic to separate these two species is that the ovaries and capsules of E. alta are distinctly papillate, with the papillae rising sharply above the surface, whereas the ovaries and capsules of E. spathulata are merely verrucose, with the protuberances lower and rounded. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
The vegetative stems of Euphorbia ipecacuanhae are often quite short in proportion to the dichasial or pleiochasial branches, thus superficially plants often appear to have mostly opposite leaves. However, careful examination of the base of the plant will reveal alternate leaves. The leaves are extremely variable in both shape and coloration, and the variation can be pronounced within a population or even on a single plant. M. J. Huft (1979) did not recognize infraspecific taxa within E. ipecacuanhae, and his treatment is followed here. This species is found on the Atlantic coastal plain. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 12, p. 298. | FNA vol. 12, p. 247. |
Parent taxa | Euphorbiaceae > Euphorbia > subg. Esula | Euphorbiaceae > Euphorbia > sect. Alectoroctonum |
Sibling taxa | ||
Synonyms | Tithymalus altus | E. arundelana, Tithymalopsis ipecacuanhae |
Name authority | Norton: N. Amer. Euphorbia, 24, plate 24. (1899) | Linnaeus: Sp. Pl. 1: 455. (1753) |
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