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rattlesnake sandmat, rattlesnake weed, white-margin sandmat or sandwort, whitemargin sandmat

eyebane, eyebane sandmat, eyebane spurge, nodding or upright spotted spurge, nodding spurge

Habit Herbs, perennial, with moderately to strongly thickened rootstock. Herbs, annual, with taproot.
Stems

prostrate, occasionally mat-forming, frequently rooting at nodes, 10–80 cm, glabrous.

usually ascending, occasionally erect, often arched at tips, 20–80 cm, sparsely to moderately pilose to villous or with short, incurved hairs, hairs often concentrated at nodes and distally, occasionally in 2 bands along opposite sides of stem.

Leaves

opposite;

stipules connate into conspicuous, deltate or ovate scale, white, 0.4–1(–2) mm, glabrous;

petiole less than 1 mm, glabrous;

blade ovate, oblong or orbiculate, 3–8(–15) × 3–7 mm, base strongly asymmetric, obtuse to hemicordate, margins whitish, entire, apex obtuse, rarely mucronulate, surfaces often with red blotch in center, glabrous; 3-veined from base but usually only midvein conspicuous.

opposite;

stipules usually distinct, sometimes connate basally on one side of stem, small scales, irregularly toothed, fringed, or divided, 1–1.5 mm, sparsely to moderately villous distally;

petiole 0.3–1.6 mm, moderately pilose to villous;

blade oblong to oblong-lanceolate, 8–40 × 3–12 mm, base asymmetric, one side usually angled or rounded, other side ± truncate to cordate-auriculate, margins serrulate, apex angled with blunt tip, abaxial surface pale green or faintly to strongly reddish tinged, adaxial surface usually reddish-mottled or with conspicuous reddish spot, both surfaces usually sparsely to moderately pilose, especially toward base, sometimes glabrous; 3–5-veined from base, pinnate distally, veins faint.

Involucre

campanulate, 0.8–1.1 × 0.9–2 mm, glabrous;

glands 4, greenish yellow to red, usually oblong to reniform, rarely subcircular, 0.2–0.5 × (0.2–)0.3–0.8 mm;

appendages white to pink, flabellate to oblong, 0.3–1 × 0.6–1.3 mm, distal margin entire or crenulate to erose.

narrowly obconic, 0.5–1 × 0.3–0.7 mm, glabrous;

glands 4, usually green, sometimes reddish purple, oblong to nearly circular, 0.2–0.4 × 0.3–0.5 mm;

appendages white or pinkish, ovate to broadly elliptic, 0.2–1 × 0.2–1.5 mm, distal margin entire.

Staminate flowers

15–30.

5–28.

Pistillate flowers

ovary glabrous;

styles 0.3–0.7 mm, 2-fid nearly entire length.

ovary glabrous;

styles 0.6–2.5 mm, 2-fid 1/2 length.

Capsules

broadly ovoid, 1.1–2.3 × 1.2–2 mm, glabrous;

columella 1.1–1.6 mm.

ovoid, 1.6–2.3 × 1.5–2.4 mm, glabrous;

columella 1.4–1.6 mm.

Seeds

white to gray or brownish red, oblong-ovoid, 4-angled in cross section, 1–1.7 × 0.5–0.8 mm, smooth.

dark brown, sometimes with thin, white coating (often more persistent along angles than faces), elliptic-ovoid to ovoid, rounded-angular in cross section, 1–1.6 × 0.5–0.8 mm, surface finely and irregularly wrinkled, sometimes faintly so, or with indistinct, shallow, rounded cross ridges.

Cyathia

solitary at distal nodes;

peduncle 1–4 mm.

solitary at distal nodes or in small, cymose clusters at branch tips;

peduncle 0.5–2.5 mm.

2n

= 12, 14, 22.

Euphorbia albomarginata

Euphorbia nutans

Phenology Flowering and fruiting year-round. Flowering and fruiting spring–early fall.
Habitat Disturbed areas in desert scrub, grasslands, mesquite woodlands, chaparral. Stream banks, pond edges, disturbed portions of upland prairies, mesic to dry upland forest openings, pastures, fallow fields, railroads, roadsides, gardens, disturbed areas.
Elevation 0–2300 m. (0–7500 ft.) 0–1600 m. (0–5200 ft.)
Distribution
from FNA
AZ; CA; CO; NM; NV; OK; TX; UT; Mexico [Introduced in Pacific Islands (Hawaii)]
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; AR; CA; CT; DE; FL; GA; IA; IL; IN; KS; KY; LA; MA; MD; MI; MN; MO; MS; NC; ND; NE; NH; NJ; NM; NY; OH; OK; PA; RI; SC; SD; TN; TX; VA; VT; WI; WV; WY; ON; QC; Mexico; Central America; South America; West Indies [Introduced in Eurasia]
[WildflowerSearch map]
[BONAP county map]
Discussion

Euphorbia albomarginata is native to northern and central Mexico and the southwestern and south-central United States. The species occurs in a variety of habitats in western North America and in some areas is quite weedy. It has been recorded as a waif in Louisiana.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Euphorbia nutans is probably native to at least central and eastern North America, but given its strongly weedy tendencies, it is difficult to know where it may be adventive in parts of the flora area. It is certainly introduced where it occurs in the Old World and probably in South America as well. D. G. Burch (1966) discussed the problems of assigning names to the four main entities in this nomenclatural complex (E. hypericifolia, E. hyssopifolia, E. lasiocarpa, and E. nutans) and determined that the oldest valid name for the relatively robust, temperate North American plants with ascending stems is E. nutans. See the treatment of 56. E. maculata for a discussion of the misapplication of that name to E. nutans.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 12, p. 259. FNA vol. 12, p. 278.
Parent taxa Euphorbiaceae > Euphorbia > sect. Anisophyllum Euphorbiaceae > Euphorbia > sect. Anisophyllum
Sibling taxa
E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana
E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana
Synonyms Chamaesyce albomarginata Chamaesyce lansingii, C. nutans, E. lansingii, E. preslii
Name authority Torrey & A. Gray: in War Department [U.S.], Pacif. Railr. Rep. 2(4): 174. (1857) Lagasca: Gen. Sp. Pl., 17. (1816)
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