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rattlesnake sandmat, rattlesnake weed, white-margin sandmat or sandwort, whitemargin sandmat

Laredo sandmat

Habit Herbs, perennial, with moderately to strongly thickened rootstock. Herbs, annual, with taproot.
Stems

prostrate, occasionally mat-forming, frequently rooting at nodes, 10–80 cm, glabrous.

prostrate, ± mat-forming, 10–20 cm, densely ashy pilose-tomentose.

Leaves

opposite;

stipules connate into conspicuous, deltate or ovate scale, white, 0.4–1(–2) mm, glabrous;

petiole less than 1 mm, glabrous;

blade ovate, oblong or orbiculate, 3–8(–15) × 3–7 mm, base strongly asymmetric, obtuse to hemicordate, margins whitish, entire, apex obtuse, rarely mucronulate, surfaces often with red blotch in center, glabrous; 3-veined from base but usually only midvein conspicuous.

opposite;

stipules distinct, filiform, 0.5–1 mm, pilose-tomentose;

petiole 0.5–1 mm, pilose-tomentose;

blade ovate to elliptic-oblong, 3–6 × 3–5 mm, base markedly asymmetric, rounded to slightly auriculate, margins usually entire, rarely largest leaves sparsely serrulate, apex acute to obtuse, surfaces moderately to densely strigose; 3-veined from base.

Involucre

campanulate, 0.8–1.1 × 0.9–2 mm, glabrous;

glands 4, greenish yellow to red, usually oblong to reniform, rarely subcircular, 0.2–0.5 × (0.2–)0.3–0.8 mm;

appendages white to pink, flabellate to oblong, 0.3–1 × 0.6–1.3 mm, distal margin entire or crenulate to erose.

obconic, 0.6–1 × 0.5–1 mm, densely strigose;

glands 4, yellowish to reddish, oval to oblong, 0.1 × 0.2–0.3 mm;

appendages white to pink, rudimentary or minute, (0–)0.1–0.2 × (0–)0.1–0.3 mm, distal margin crenulate.

Staminate flowers

15–30.

3–5.

Pistillate flowers

ovary glabrous;

styles 0.3–0.7 mm, 2-fid nearly entire length.

ovary densely white villous;

styles 0.1–0.2 mm, 2-fid 1/2 length.

Capsules

broadly ovoid, 1.1–2.3 × 1.2–2 mm, glabrous;

columella 1.1–1.6 mm.

broadly ovoid, 1.3–1.5 × 1.4–1.5 mm, villous on keels, often glabrous or less hairy between keels;

columella 1.1–1.3 mm.

Seeds

white to gray or brownish red, oblong-ovoid, 4-angled in cross section, 1–1.7 × 0.5–0.8 mm, smooth.

white, barely concealing brown undercoat, 4-angled, sharply angled in cross section, abaxial faces plane to convex, adaxial faces concave, 1.1–1.2 × 0.5–0.7 mm, with several rounded, irregular, transverse ridges.

Cyathia

solitary at distal nodes;

peduncle 1–4 mm.

solitary or in small, cymose clusters at distal nodes or on congested, axillary branches;

peduncle 0.5–1.5 mm.

Euphorbia albomarginata

Euphorbia laredana

Phenology Flowering and fruiting year-round. Flowering and fruiting almost year-round.
Habitat Disturbed areas in desert scrub, grasslands, mesquite woodlands, chaparral. Open sandy, loamy, or gravelly sites, old dunes, pastures.
Elevation 0–2300 m. (0–7500 ft.) 0–200 m. (0–700 ft.)
Distribution
from FNA
AZ; CA; CO; NM; NV; OK; TX; UT; Mexico [Introduced in Pacific Islands (Hawaii)]
[WildflowerSearch map]
[BONAP county map]
from FNA
TX; Mexico (Tamaulipas)
[BONAP county map]
Discussion

Euphorbia albomarginata is native to northern and central Mexico and the southwestern and south-central United States. The species occurs in a variety of habitats in western North America and in some areas is quite weedy. It has been recorded as a waif in Louisiana.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Euphorbia laredana is similar to E. prostrata but differs from that species in its more densely tomentose indumentum, leaves with usually entire rather than serrulate margins, and slightly longer seeds with rounded rather than sharp ridges. The species occurs primarily in southern Texas.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 12, p. 259. FNA vol. 12, p. 274.
Parent taxa Euphorbiaceae > Euphorbia > sect. Anisophyllum Euphorbiaceae > Euphorbia > sect. Anisophyllum
Sibling taxa
E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana
E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana
Synonyms Chamaesyce albomarginata Chamaesyce laredana
Name authority Torrey & A. Gray: in War Department [U.S.], Pacif. Railr. Rep. 2(4): 174. (1857) Millspaugh: Pittonia 2: 88. (1890)
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