Euphorbia albomarginata |
Euphorbia hooveri |
|
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rattlesnake sandmat, rattlesnake weed, white-margin sandmat or sandwort, whitemargin sandmat |
Hoover's sandmat, Hoover's spurge |
|
Habit | Herbs, perennial, with moderately to strongly thickened rootstock. | Herbs, annual, with slender to slightly thickened taproot. |
Stems | prostrate, occasionally mat-forming, frequently rooting at nodes, 10–80 cm, glabrous. |
prostrate, occasionally mat-forming, 4–10 cm, glabrous. |
Leaves | opposite; stipules connate into conspicuous, deltate or ovate scale, white, 0.4–1(–2) mm, glabrous; petiole less than 1 mm, glabrous; blade ovate, oblong or orbiculate, 3–8(–15) × 3–7 mm, base strongly asymmetric, obtuse to hemicordate, margins whitish, entire, apex obtuse, rarely mucronulate, surfaces often with red blotch in center, glabrous; 3-veined from base but usually only midvein conspicuous. |
opposite; stipules distinct or connate at base, deeply laciniate into numerous subulate to filiform segments, 0.3–1.3 mm, glabrous; petiole 0–0.5 mm, glabrous; blade broadly ovate, broadly oblong or suborbiculate, 3–7 × 2–5.5 mm, base asymmetric, hemiamplexicaulous, margin sharply serrulate to denticulate, teeth whitish and occasionally setaelike at apex, apex obtuse, surfaces papillate, glabrous; 3-veined from base, usually only midvein conspicuous. |
Involucre | campanulate, 0.8–1.1 × 0.9–2 mm, glabrous; glands 4, greenish yellow to red, usually oblong to reniform, rarely subcircular, 0.2–0.5 × (0.2–)0.3–0.8 mm; appendages white to pink, flabellate to oblong, 0.3–1 × 0.6–1.3 mm, distal margin entire or crenulate to erose. |
campanulate, 1.2–1.5 × 1.5–2.2 mm, papillate; glands 4, yellow to reddish, subcircular to oblong, 0.4–0.5 × 0.5–0.7 mm; appendages white to pink, 0.3–0.9 × 0.7–1.6 mm, divided into 4–6 narrowly triangular, acute segments, these occasionally 2-fid, margins entire. |
Staminate flowers | 15–30. |
25–35. |
Pistillate flowers | ovary glabrous; styles 0.3–0.7 mm, 2-fid nearly entire length. |
ovary glabrous; styles 1.8–2.6 mm, unbranched, filiform. |
Capsules | broadly ovoid, 1.1–2.3 × 1.2–2 mm, glabrous; columella 1.1–1.6 mm. |
oblate, 1.5–1.7 × 1.8–2.2 mm, glabrous; columella 1.2–1.4 mm. |
Seeds | white to gray or brownish red, oblong-ovoid, 4-angled in cross section, 1–1.7 × 0.5–0.8 mm, smooth. |
gray to light brown, ovoid, inconspicuously 4-angled in cross section, 1.2–1.6 × 1–1.1 mm, rugose. |
Cyathia | solitary at distal nodes; peduncle 1–4 mm. |
solitary at distal nodes; peduncle 0.5 mm, stout. |
Euphorbia albomarginata |
Euphorbia hooveri |
|
Phenology | Flowering and fruiting year-round. | Flowering and fruiting summer–fall. |
Habitat | Disturbed areas in desert scrub, grasslands, mesquite woodlands, chaparral. | Drying mudflats of vernal pools in grasslands and woodlands. |
Elevation | 0–2300 m. (0–7500 ft.) | 20–200 m. (100–700 ft.) |
Distribution |
AZ; CA; CO; NM; NV; OK; TX; UT; Mexico [Introduced in Pacific Islands (Hawaii)]
|
CA |
Discussion | Euphorbia albomarginata is native to northern and central Mexico and the southwestern and south-central United States. The species occurs in a variety of habitats in western North America and in some areas is quite weedy. It has been recorded as a waif in Louisiana. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Euphorbia hooveri is federally listed as threatened; its populations are being affected severely by habitat loss and the invasion of exotic species. The species is endemic to vernal pools in six counties in the Central Valley of California. Molecular data show that E. hooveri is a hybrid species, closely related to E. albomarginata and E. serpens (Y. Yang and P. E. Berry 2011). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 12, p. 259. | FNA vol. 12, p. 271. |
Parent taxa | Euphorbiaceae > Euphorbia > sect. Anisophyllum | Euphorbiaceae > Euphorbia > sect. Anisophyllum |
Sibling taxa | ||
Synonyms | Chamaesyce albomarginata | Chamaesyce hooveri |
Name authority | Torrey & A. Gray: in War Department [U.S.], Pacif. Railr. Rep. 2(4): 174. (1857) | L. C. Wheeler: Proc. Biol. Soc. Wash. 53: 9. (1940) |
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