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rattlesnake sandmat, rattlesnake weed, white-margin sandmat or sandwort, whitemargin sandmat

Hoover's sandmat, Hoover's spurge

Habit Herbs, perennial, with moderately to strongly thickened rootstock. Herbs, annual, with slender to slightly thickened taproot.
Stems

prostrate, occasionally mat-forming, frequently rooting at nodes, 10–80 cm, glabrous.

prostrate, occasionally mat-forming, 4–10 cm, glabrous.

Leaves

opposite;

stipules connate into conspicuous, deltate or ovate scale, white, 0.4–1(–2) mm, glabrous;

petiole less than 1 mm, glabrous;

blade ovate, oblong or orbiculate, 3–8(–15) × 3–7 mm, base strongly asymmetric, obtuse to hemicordate, margins whitish, entire, apex obtuse, rarely mucronulate, surfaces often with red blotch in center, glabrous; 3-veined from base but usually only midvein conspicuous.

opposite;

stipules distinct or connate at base, deeply laciniate into numerous subulate to filiform segments, 0.3–1.3 mm, glabrous;

petiole 0–0.5 mm, glabrous;

blade broadly ovate, broadly oblong or suborbiculate, 3–7 × 2–5.5 mm, base asymmetric, hemiamplexicaulous, margin sharply serrulate to denticulate, teeth whitish and occasionally setaelike at apex, apex obtuse, surfaces papillate, glabrous; 3-veined from base, usually only midvein conspicuous.

Involucre

campanulate, 0.8–1.1 × 0.9–2 mm, glabrous;

glands 4, greenish yellow to red, usually oblong to reniform, rarely subcircular, 0.2–0.5 × (0.2–)0.3–0.8 mm;

appendages white to pink, flabellate to oblong, 0.3–1 × 0.6–1.3 mm, distal margin entire or crenulate to erose.

campanulate, 1.2–1.5 × 1.5–2.2 mm, papillate;

glands 4, yellow to reddish, subcircular to oblong, 0.4–0.5 × 0.5–0.7 mm;

appendages white to pink, 0.3–0.9 × 0.7–1.6 mm, divided into 4–6 narrowly triangular, acute segments, these occasionally 2-fid, margins entire.

Staminate flowers

15–30.

25–35.

Pistillate flowers

ovary glabrous;

styles 0.3–0.7 mm, 2-fid nearly entire length.

ovary glabrous;

styles 1.8–2.6 mm, unbranched, filiform.

Capsules

broadly ovoid, 1.1–2.3 × 1.2–2 mm, glabrous;

columella 1.1–1.6 mm.

oblate, 1.5–1.7 × 1.8–2.2 mm, glabrous;

columella 1.2–1.4 mm.

Seeds

white to gray or brownish red, oblong-ovoid, 4-angled in cross section, 1–1.7 × 0.5–0.8 mm, smooth.

gray to light brown, ovoid, inconspicuously 4-angled in cross section, 1.2–1.6 × 1–1.1 mm, rugose.

Cyathia

solitary at distal nodes;

peduncle 1–4 mm.

solitary at distal nodes;

peduncle 0.5 mm, stout.

Euphorbia albomarginata

Euphorbia hooveri

Phenology Flowering and fruiting year-round. Flowering and fruiting summer–fall.
Habitat Disturbed areas in desert scrub, grasslands, mesquite woodlands, chaparral. Drying mudflats of vernal pools in grasslands and woodlands.
Elevation 0–2300 m. (0–7500 ft.) 20–200 m. (100–700 ft.)
Distribution
from FNA
AZ; CA; CO; NM; NV; OK; TX; UT; Mexico [Introduced in Pacific Islands (Hawaii)]
[WildflowerSearch map]
[BONAP county map]
from FNA
CA
[BONAP county map]
Discussion

Euphorbia albomarginata is native to northern and central Mexico and the southwestern and south-central United States. The species occurs in a variety of habitats in western North America and in some areas is quite weedy. It has been recorded as a waif in Louisiana.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Euphorbia hooveri is federally listed as threatened; its populations are being affected severely by habitat loss and the invasion of exotic species. The species is endemic to vernal pools in six counties in the Central Valley of California. Molecular data show that E. hooveri is a hybrid species, closely related to E. albomarginata and E. serpens (Y. Yang and P. E. Berry 2011).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 12, p. 259. FNA vol. 12, p. 271.
Parent taxa Euphorbiaceae > Euphorbia > sect. Anisophyllum Euphorbiaceae > Euphorbia > sect. Anisophyllum
Sibling taxa
E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana
E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana
Synonyms Chamaesyce albomarginata Chamaesyce hooveri
Name authority Torrey & A. Gray: in War Department [U.S.], Pacif. Railr. Rep. 2(4): 174. (1857) L. C. Wheeler: Proc. Biol. Soc. Wash. 53: 9. (1940)
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