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rattlesnake sandmat, rattlesnake weed, white-margin sandmat or sandwort, whitemargin sandmat

wedge sandmat

Habit Herbs, perennial, with moderately to strongly thickened rootstock. Herbs, perennial, delicate, with woody, thickened taproot, 15 mm diam..
Stems

prostrate, occasionally mat-forming, frequently rooting at nodes, 10–80 cm, glabrous.

prostrate, ascending, or erect, often numerous and wiry, less than 0.1 mm diam., 5–20 cm, glabrous, puberulent, canescent, villous, or hirsute, shorter hairs often uncinate and longer hairs straight or irregularly twisted.

Leaves

opposite;

stipules connate into conspicuous, deltate or ovate scale, white, 0.4–1(–2) mm, glabrous;

petiole less than 1 mm, glabrous;

blade ovate, oblong or orbiculate, 3–8(–15) × 3–7 mm, base strongly asymmetric, obtuse to hemicordate, margins whitish, entire, apex obtuse, rarely mucronulate, surfaces often with red blotch in center, glabrous; 3-veined from base but usually only midvein conspicuous.

opposite;

stipules distinct, triangular, sometimes lacerate or ciliate, 0.2–0.3 mm, glabrous or hairy;

petiole 0.3–1 mm, glabrous or hairy;

blade narrowly to broadly deltate, cordate, or reniform, 2–5(–7) × 1–4.5(–5) mm, base asymmetric, cordate to rounded, margins entire, ± revolute, apex obtuse or rounded, surfaces glabrous or hairy;

only midvein conspicuous.

Involucre

campanulate, 0.8–1.1 × 0.9–2 mm, glabrous;

glands 4, greenish yellow to red, usually oblong to reniform, rarely subcircular, 0.2–0.5 × (0.2–)0.3–0.8 mm;

appendages white to pink, flabellate to oblong, 0.3–1 × 0.6–1.3 mm, distal margin entire or crenulate to erose.

turbinate to campanulate, 0.8–1 × 1.1–1.3 mm, glabrous or hairy;

glands 4, green to yellow-green, oblong to subcircular, 0.2–0.4 × 0.4–0.6 mm;

appendages absent or white, forming narrow rim at edge of gland, rarely slightly wider than gland, (0–)0.1(–0.3) × 0.4–0.6 mm, distal margin entire.

Staminate flowers

15–30.

8–14.

Pistillate flowers

ovary glabrous;

styles 0.3–0.7 mm, 2-fid nearly entire length.

ovary glabrous or hairy, subtended by triangular pad of tissue;

styles spreading, 0.3–0.4 mm, 2-fid 1/2 to nearly entire length.

Capsules

broadly ovoid, 1.1–2.3 × 1.2–2 mm, glabrous;

columella 1.1–1.6 mm.

broadly deltoid, 1.2–1.5 × 2–2.2 mm, glabrous or hairy;

columella 0.9–1.3 mm.

Seeds

white to gray or brownish red, oblong-ovoid, 4-angled in cross section, 1–1.7 × 0.5–0.8 mm, smooth.

reddish brown, ovoid, 4-angled in cross section, 0.8–1.2 × 0.5–0.6 mm, obscurely wrinkled.

Cyathia

solitary at distal nodes;

peduncle 1–4 mm.

solitary at distal nodes;

peduncle 0.7–1.5 mm.

Euphorbia albomarginata

Euphorbia deltoidea

Phenology Flowering and fruiting year-round.
Habitat Disturbed areas in desert scrub, grasslands, mesquite woodlands, chaparral.
Elevation 0–2300 m. (0–7500 ft.)
Distribution
from FNA
AZ; CA; CO; NM; NV; OK; TX; UT; Mexico [Introduced in Pacific Islands (Hawaii)]
[WildflowerSearch map]
[BONAP county map]
from FNA
FL
[BONAP county map]
Discussion

Euphorbia albomarginata is native to northern and central Mexico and the southwestern and south-central United States. The species occurs in a variety of habitats in western North America and in some areas is quite weedy. It has been recorded as a waif in Louisiana.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Subspecies 4 (4 in the flora).

Euphorbia deltoidea comprises four narrowly endemic subspecies, all of which are endangered due to habitat loss and fragmentation. The subspecies occur in pine rockland habitat that is free of shrubby undergrowth. Periodic fires are required to keep the rockland habitat open. Subspecies serpyllum is restricted to Big Pine Key, Monroe County, whereas the other subspecies are found only in Miami-Dade County.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Stems erect or ascending.
→ 2
2. Leaves and stems villous, hairs uncinate or irregularly twisted, 0.2–0.5 mm; leaf blades green.
subsp. adhaerens
2. Leaves and stems villous-hirsute, hairs straight and spreading, 0.6–0.7 mm; leaf blades silver-green.
subsp. pinetorum
1. Stems prostrate.
→ 3
3. Leaves and stems glabrous or very sparsely hairy, hairs 0.1–0.2 mm, appressed, uncinate; leaf blades abaxially reddish, adaxially bright green.
subsp. deltoidea
3. Leaves and stems canescent or villous, hairs either less than 0.1 mm or 0.2–0.5 mm, uncinate or irregularly twisted; leaf blades green or silver-green.
→ 4
4. Leaves and stems villous, hairs uncinate or irregularly twisted, 0.2–0.5 mm; leaf blades as long as wide, green.
subsp. adhaerens
4. Leaves and stems canescent, hairs less than 0.1 mm; leaf blades 2 times longer than wide, silver-green.
subsp. serpyllum
Source FNA vol. 12, p. 259. FNA vol. 12, p. 265.
Parent taxa Euphorbiaceae > Euphorbia > sect. Anisophyllum Euphorbiaceae > Euphorbia > sect. Anisophyllum
Sibling taxa
E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana
E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana
Subordinate taxa
E. deltoidea subsp. adhaerens, E. deltoidea subsp. deltoidea, E. deltoidea subsp. pinetorum, E. deltoidea subsp. serpyllum
Synonyms Chamaesyce albomarginata Chamaesyce deltoidea
Name authority Torrey & A. Gray: in War Department [U.S.], Pacif. Railr. Rep. 2(4): 174. (1857) Engelmann ex Chapman: Fl. South. U.S. ed. 2, 647. (1883)
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