Euphorbia agraria |
Euphorbia villifera |
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urban spurge |
hairy spurge |
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Habit | Herbs, perennial, with slender, spreading rootstock. | Herbs, annual or perennial, with slender taproot or thickened, woody rootstock. |
Stems | erect or ascending, unbranched or branched, 30–90 cm, glabrous. |
usually erect to ascending, rarely prostrate to decumbent, 10–30 cm, papillate, usually villous, sometimes glabrous. |
Leaves | petiole absent; blade oblong-elliptic, 20–65 × 9–20 mm, base truncate to auriculate, margins entire, apex obtuse to rounded, surfaces glabrous; venation conspicuously pinnate, midvein prominent. |
opposite; stipules distinct, filiform, usually undivided, rarely divided into 2–3 segments), 0.3–0.7 mm, glabrous, papillate; petiole 0.6–1.8 mm, usually villous, rarely glabrous; blade ovate, 3–12 × 2–10 mm, base asymmetric, rounded to slightly cordate, margins entire or serrulate, apex acute to obtuse, surfaces usually villous, rarely glabrous; only midvein conspicuous. |
Involucre | campanulate, 2.2–3 × 1.8–2 mm, glabrous; glands 4, crescent-shaped; 0.6–1 × 1–2 mm; horns slightly divergent to convergent, 0.1–0.2 mm. |
campanulate, 0.7–0.9 × 0.6–1 mm, glabrous or pilose; glands 4, pink, oval, oblong, or trapezoidal, 0.1–0.2 × 0.2 mm; appendages white to pink, flabellate, oblong, ovate, or nearly rectangular, 0.2–0.4 × 0.2–0.6 mm, distal margin entire. |
Staminate flowers | 15–20. |
10–25. |
Pistillate flowers | ovary glabrous; styles 1.2–2 mm, 2-fid. |
ovary glabrous; styles 0.3–0.5 mm, 2-fid 1/2 to nearly entire length. |
Capsules | globose, 2–2.8 × 2.2–2.7 mm, 3-lobed; cocci rounded, smooth except finely granulate toward abaxial line, glabrous; columella 2.1–2.7 mm. |
oblate-deltoid, cocci often elongated and terminating in an empty portion, 1.5–2 × 2.1–3.1 mm, glabrous; columella 0.9–1.5 mm. |
Seeds | gray or whitish, ovoid-oblong, 2–2.1 × 1.2–1.3 mm, smooth; caruncle ± rounded and flattened, 0.8 × 0.6 mm. |
gray-brown to red-brown, ovoid-oblong, weakly 4-angled in cross section, 1–1.4 × 0.6–0.8 mm, smooth, faintly rugose, or with inconspicuous transverse ridges. |
Cyathial | arrangement: terminal pleiochasial branches 8–15, 1–2 times 2-branched; pleiochasial bracts similar in shape but shorter and narrower than distal leaves; dichasial bracts distinct, rhombic to reniform, base obtuse, margins entire, apex obtuse, mucronate; axillary cymose branches 12–23. |
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Cyathia | peduncle 0–2 mm. |
solitary at distal nodes; peduncles 0–1.8 mm. |
Euphorbia agraria |
Euphorbia villifera |
|
Phenology | Flowering and fruiting spring–summer. | Flowering and fruiting early spring–early winter. |
Habitat | Grasslands, roadside banks, pastures. | Riparian forests with walnuts and sycamores, juniper woodlands, pine-oak woodlands, mostly on limestone substrates. |
Elevation | 200–1600 m. (700–5200 ft.) | 100–1400 m. (300–4600 ft.) |
Distribution |
KS; MT; NE; NY; PA; WA; WY; AB; SK; Europe [Introduced in North America]
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TX; Mexico; Central America |
Discussion | Although Euphorbia villifera has been reported from New Mexico (W. C. Martin and C. R. Hutchins 1980), no vouchers to verify its presence there were located. In Texas, E. villifera is known from the Edwards Plateau westward into the trans-Pecos region. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
|
Source | FNA vol. 12, p. 298. | FNA vol. 12, p. 293. |
Parent taxa | Euphorbiaceae > Euphorbia > subg. Esula | Euphorbiaceae > Euphorbia > sect. Anisophyllum |
Sibling taxa | ||
Synonyms | Tithymalus agrarius | Chamaesyce stanfieldii, C. villifera, E. stanfieldii, E. villifera var. nuda |
Name authority | M. Bieberstein: Fl. Taur.-Caucas. 1: 375. (1808) | Scheele: Linnaea 22: 153. (1849) |
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