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urban spurge

Caliche sandmat

Habit Herbs, perennial, with slender, spreading rootstock. Herbs, perennial, with strongly thickened and lignified rootstock.
Stems

erect or ascending, unbranched or branched, 30–90 cm, glabrous.

prostrate, usually mat-forming, terete to slightly flattened and winged, to 30 cm, villous to pilose.

Leaves

petiole absent;

blade oblong-elliptic, 20–65 × 9–20 mm, base truncate to auriculate, margins entire, apex obtuse to rounded, surfaces glabrous;

venation conspicuously pinnate, midvein prominent.

opposite;

stipules distinct or connate at base, 2-fid to laciniate into 2–5 linear to subulate divisions, 0.5–1(–2) mm, glabrous or villous;

petiole 0.5–2 mm, villous;

blade usually ovate to oblong, rarely suborbiculate, 5–13 × 4–8 mm, base asymmetric, hemicordate, margins serrulate, apex obtuse, surfaces villous; 3-veined from base, usually only midvein conspicuous.

Involucre

campanulate, 2.2–3 × 1.8–2 mm, glabrous;

glands 4, crescent-shaped; 0.6–1 × 1–2 mm;

horns slightly divergent to convergent, 0.1–0.2 mm.

turbinate, 0.9–1.4 × 0.7–1.1 mm, villous;

glands 4, pink, oblong to reniform, 0.1–0.2 × 0.3–0.4 mm;

appendages white to pink, flabellate, ovate, or oblong, 0.3–0.5 × 0.4–0.7 mm, distal margin entire, crenulate, or erose, often puberulent-ciliate.

Staminate flowers

15–20.

8–12.

Pistillate flowers

ovary glabrous;

styles 1.2–2 mm, 2-fid.

ovary villous;

styles 0.2–0.4(–0.5) mm, 2-fid 1/2 length.

Capsules

globose, 2–2.8 × 2.2–2.7 mm, 3-lobed;

cocci rounded, smooth except finely granulate toward abaxial line, glabrous;

columella 2.1–2.7 mm.

broadly ovoid, 1.5–1.9 × 1.3–1.7 mm, villous (uniformly so, but most pronounced toward base and along keels);

columella 1.2–1.4 mm.

Seeds

gray or whitish, ovoid-oblong, 2–2.1 × 1.2–1.3 mm, smooth;

caruncle ± rounded and flattened, 0.8 × 0.6 mm.

white to gray or pink, narrowly pyramidal-ovoid, 4-angled in cross section, 1.1–1.3 × 0.4–0.6 mm, rugulose with shallow depressions separated by inconspicuous transverse ridges.

Cyathial

arrangement: terminal pleiochasial branches 8–15, 1–2 times 2-branched;

pleiochasial bracts similar in shape but shorter and narrower than distal leaves;

dichasial bracts distinct, rhombic to reniform, base obtuse, margins entire, apex obtuse, mucronate;

axillary cymose branches 12–23.

Cyathia

peduncle 0–2 mm.

solitary at distal nodes or at nodes of leafy, congested, axillary branches;

peduncle 1–1.5(–2.5) mm.

Euphorbia agraria

Euphorbia velleriflora

Phenology Flowering and fruiting spring–summer. Flowering and fruiting summer.
Habitat Grasslands, roadside banks, pastures. Disturbed habitats, particularly roadsides, grasslands, live oak-thorn scrub.
Elevation 200–1600 m. (700–5200 ft.) 50–80 m. (200–300 ft.)
Distribution
from FNA
KS; MT; NE; NY; PA; WA; WY; AB; SK; Europe [Introduced in North America]
[WildflowerSearch map]
[BONAP county map]
from FNA
TX; Mexico; Central America (Guatemala) [Introduced in North America]
[BONAP county map]
Discussion

Euphorbia velleriflora is native and widespread from northern Mexico to Guatemala. The species is apparently a recent introduction in southern Texas (W. R. Carr and M. H. Mayfield 1993), as the first records are from the 1990s. It is also expected in southern Arizona due to the presence of collections made a few kilometers south of the border in the Mexican state of Sonora. Euphorbia velleriflora is very similar to E. stictospora, and the two are sometimes confused. The two can be readily distinguished on the basis of their styles: unbranched in E. stictospora and 2-fid in E. velleriflora. Also, the involucral gland appendages of E. velleriflora are ciliate with short hairs, whereas those of E. stictospora are glabrous.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 12, p. 298. FNA vol. 12, p. 292.
Parent taxa Euphorbiaceae > Euphorbia > subg. Esula Euphorbiaceae > Euphorbia > sect. Anisophyllum
Sibling taxa
E. aaron-rossii, E. abramsiana, E. acuta, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana
E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana
Synonyms Tithymalus agrarius Anisophyllum velleriflorum, Chamaesyce velleriflora
Name authority M. Bieberstein: Fl. Taur.-Caucas. 1: 375. (1808) (Klotzsch & Garcke) Boissier: in A. P. de Candolle and A. L. P. P. de Candolle, Prodr. 15(2): 40. (1862)
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