Flora of North America species comparison

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Euphorbia agraria

Euphorbia geyeri

urban spurge

Geyer's sandmat, Geyer's spurge
Habit Herbs, perennial, with slender, spreading rootstock. Herbs, annual, with taproot.
Stems

erect or ascending, unbranched or branched, 30–90 cm, glabrous.

prostrate or slightly ascending, loosely mat-forming, 4–25(–45) cm, glabrous.
Leaves

petiole absent;

blade oblong-elliptic, 20–65 × 9–20 mm, base truncate to auriculate, margins entire, apex obtuse to rounded, surfaces glabrous;

venation conspicuously pinnate, midvein prominent.

opposite;

stipules usually distinct, occasionally connate basally on lower side, usually deeply parted into 3 or more filiform segments, 0.7–1.5 mm, glabrous;

petiole 1–2 mm, glabrous;

blade oblong to oblong-obovate or oblong-elliptic, 4–12 × 2–6 mm, base slightly asymmetric, angled or rounded, with one side usually expanded into small, rounded auricle, margins entire, apex usually truncate, occasionally emarginate, abaxial surface pale grayish green, both surfaces glabrous;

only midvein conspicuous or venation obscurely pinnate (larger leaves).
Involucre

campanulate, 2.2–3 × 1.8–2 mm, glabrous;

glands 4, crescent-shaped; 0.6–1 × 1–2 mm;

horns slightly divergent to convergent, 0.1–0.2 mm.

broadly campanulate, 1–1.5 × 0.7–0.9 mm, glabrous;

glands 4, green to reddish, elliptic-oblong to nearly circular, slightly cupped to folded, 0.2–0.4 × 0.2–0.6 mm;

appendages rudimentary to absent or white to reddish-tinged, usually rounded, sometimes pointed, (0–)0.5–1 × (0–)0.1–1.2 mm, distal margin entire or slightly toothed.
Staminate flowers

15–20.

5–20.
Pistillate flowers

ovary glabrous;

styles 1.2–2 mm, 2-fid.

ovary glabrous;

styles 0.2–0.6 mm, 2-fid nearly 1/2 length.
Capsules

globose, 2–2.8 × 2.2–2.7 mm, 3-lobed;

cocci rounded, smooth except finely granulate toward abaxial line, glabrous;

columella 2.1–2.7 mm.

globose-ovoid, 1.5–2 × 1.5–3 mm, glabrous;

columella 1.5–1.9 mm.
Seeds

gray or whitish, ovoid-oblong, 2–2.1 × 1.2–1.3 mm, smooth;

caruncle ± rounded and flattened, 0.8 × 0.6 mm.

ashy white, ovoid, terete to bluntly subangled in cross section, 1.1–1.7 × 0.9–1.2 mm, smooth, with smooth brown line from top to bottom on adaxial side.
Cyathial

arrangement: terminal pleiochasial branches 8–15, 1–2 times 2-branched;

pleiochasial bracts similar in shape but shorter and narrower than distal leaves;

dichasial bracts distinct, rhombic to reniform, base obtuse, margins entire, apex obtuse, mucronate;

axillary cymose branches 12–23.

Cyathia

peduncle 0–2 mm.

solitary or in small, cymose clusters at distal nodes;

peduncle 1–2 mm.

Euphorbia agraria

Euphorbia geyeri
Phenology Flowering and fruiting spring–summer.
Habitat Grasslands, roadside banks, pastures.
Elevation 200–1600 m. (700–5200 ft.)
Distribution
from FNA
KS; MT; NE; NY; PA; WA; WY; AB; SK; Europe [Introduced in North America]
[WildflowerSearch map]
[BONAP county map]
from FNA
CO; IA; IL; IN; KS; LA; MI; MN; MO; MS; MT; ND; NE; NM; OK; SD; TX; UT; WI; WY; MB; n Mexico
[WildflowerSearch map]
[BONAP county map]
Discussion

Varieties 2 (2 in the flora).

The two varieties of Euphorbia geyeri have been distinguished in large part by the presence of conspicuous involucral gland appendages in var. geyeri and the lack of appendages in var. wheeleriana. The two varieties are recognized here, but the variation in the size and presence of involucral gland appendages in the closely related E. bombensis suggests that this might be a somewhat variable character in this group of species.

Euphorbia geyeri is widespread throughout the central United States in sandy soils. Populations at the eastern edge of the range are often considered adventive (for example, sandy soils along railroad grades in Michigan). Euphorbia geyeri resembles E. glyptosperma (both being entirely glabrous), but that species has serrulate leaves (near the apex) and strongly angled, transverse-ridged seeds whereas E. geyeri has entire leaves and smooth, rounded seeds.

(Discussion copyrighted by Flora of North America; reprinted with permission.)
Key
1. Involucral gland appendages present; staminate flowers 5–9; seeds 1.1–1.4(–1.6) mm.
var. geyeri
1. Involucral gland appendages absent or rudimentary; staminate flowers 10–20; seeds 1.6–1.7 mm.
var. wheeleriana
Source FNA vol. 12, p. 298. FNA vol. 12, p. 268.
Parent taxa Euphorbiaceae > Euphorbia > subg. Esula Euphorbiaceae > Euphorbia > sect. Anisophyllum
Sibling taxa
E. aaron-rossii, E. abramsiana, E. acuta, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana
E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana
Subordinate taxa
E. geyeri var. geyeri, E. geyeri var. wheeleriana
Synonyms Tithymalus agrarius Chamaesyce geyeri
Name authority M. Bieberstein: Fl. Taur.-Caucas. 1: 375. (1808) Engelmann: Boston J. Nat. Hist. 5: 260. (1845)
Web links 
Unless otherwise noted, all data comes from the online version of the Flora of North America, who have made most of their content available via a Creative Commons license (CC BY 4.0).



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