Euphorbia agraria |
Euphorbia garberi |
|
---|---|---|
urban spurge |
Garber's sandmat |
|
Habit | Herbs, perennial, with slender, spreading rootstock. | Herbs, perennial, with moderately thickened rootstock. |
Stems | erect or ascending, unbranched or branched, 30–90 cm, glabrous. |
ascending, sometimes slightly woody at base, not wiry, 1–3 mm diam., 15–50 cm, canescent. |
Leaves | petiole absent; blade oblong-elliptic, 20–65 × 9–20 mm, base truncate to auriculate, margins entire, apex obtuse to rounded, surfaces glabrous; venation conspicuously pinnate, midvein prominent. |
opposite; stipules distinct or connate at base, triangular-subulate, apex divided into 2–5 subulate to subulate-filiform segments, 0.5–0.7 mm, pilose; petiole 0.5–1.1 mm, pilose; blade ovate to oblong-elliptic, 4–9(–15) × 3–6 mm, base asymmetric, obtuse to rounded, margins usually entire, rarely very sparsely serrulate, apex usually obtuse to rounded, occasionally acute to apiculate, surfaces canescent; 3-veined from base, only midvein conspicuous. |
Involucre | campanulate, 2.2–3 × 1.8–2 mm, glabrous; glands 4, crescent-shaped; 0.6–1 × 1–2 mm; horns slightly divergent to convergent, 0.1–0.2 mm. |
turbinate to campanulate, 0.6–0.9 × 0.5–0.9 mm, pilose; glands 4, brown, slightly concave, elliptic to subcircular, 0.1–0.3 × 0.2–0.4 mm; appendages white to pink, as narrow rim along gland, (0–)0.1–0.2 × 0.3–0.5 mm, surfaces pilose, distal margin entire or crenulate. |
Staminate flowers | 15–20. |
8–20. |
Pistillate flowers | ovary glabrous; styles 1.2–2 mm, 2-fid. |
ovary pilose; styles 0.6–0.7 mm, 2-fid 1/2 length. |
Capsules | globose, 2–2.8 × 2.2–2.7 mm, 3-lobed; cocci rounded, smooth except finely granulate toward abaxial line, glabrous; columella 2.1–2.7 mm. |
subglobose to broadly ovoid, 1.1–1.6 × 1.3–2.1 mm, pilose; columella 1.1–1.6 mm. |
Seeds | gray or whitish, ovoid-oblong, 2–2.1 × 1.2–1.3 mm, smooth; caruncle ± rounded and flattened, 0.8 × 0.6 mm. |
gray to reddish brown, oblong-ovoid, 4-angled in cross section, 1–1.2 × 0.6–0.8 mm, flat or obscurely wrinkled. |
Cyathial | arrangement: terminal pleiochasial branches 8–15, 1–2 times 2-branched; pleiochasial bracts similar in shape but shorter and narrower than distal leaves; dichasial bracts distinct, rhombic to reniform, base obtuse, margins entire, apex obtuse, mucronate; axillary cymose branches 12–23. |
|
Cyathia | peduncle 0–2 mm. |
solitary at nodes of short, axillary branches; peduncle 0.2–0.4 mm. |
Euphorbia agraria |
Euphorbia garberi |
|
Phenology | Flowering and fruiting spring–summer. | Flowering and fruiting year-round. |
Habitat | Grasslands, roadside banks, pastures. | Beach dunes, coastal rock barrens, disturbed uplands and pine rocklands. |
Elevation | 200–1600 m. (700–5200 ft.) | 0–10 m. (0–0 ft.) |
Distribution |
KS; MT; NE; NY; PA; WA; WY; AB; SK; Europe [Introduced in North America]
|
FL |
Discussion | Euphorbia garberi appears to be an interspecific hybrid, but its parentage is not entirely clear (Y. Yang and P. E. Berry 2011). The taxon is closely related to E. blodgettii, E. porteriana, and E. serpens. It is restricted to Collier, Miami-Dade, and Monroe counties in southern Florida and is federally listed as threatened due to impacts from habitat loss, fire suppression, and invasive species. A. Herndon (1993) synonymized E. porteriana var. keyensis under E. garberi because of its uniformly pilose capsules, and that treatment is followed here. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
|
Source | FNA vol. 12, p. 298. | FNA vol. 12, p. 268. |
Parent taxa | Euphorbiaceae > Euphorbia > subg. Esula | Euphorbiaceae > Euphorbia > sect. Anisophyllum |
Sibling taxa | ||
Synonyms | Tithymalus agrarius | Chamaesyce adicioides, C. brachypoda, C. garberi, C. keyensis, C. mosieri, C. porteriana var. keyensis, E. porteriana var. keyensis |
Name authority | M. Bieberstein: Fl. Taur.-Caucas. 1: 375. (1808) | Engelmann ex Chapman: Fl. South. U.S. ed. 2, 646. (1883) |
Web links |