Euphorbia agraria |
Euphorbia davidii |
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urban spurge |
David's poinsettia, David's spurge, tooth poinsettia, tooth spurge |
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Habit | Herbs, perennial, with slender, spreading rootstock. | Herbs, annual, with taproot. |
Stems | erect or ascending, unbranched or branched, 30–90 cm, glabrous. |
erect or ascending, 20–70 cm, both coarsely and sparsely hirsute and closely strigillose; branches usually ± straight, occasionally proximal branches arcuate. |
Leaves | petiole absent; blade oblong-elliptic, 20–65 × 9–20 mm, base truncate to auriculate, margins entire, apex obtuse to rounded, surfaces glabrous; venation conspicuously pinnate, midvein prominent. |
usually opposite, occasionally alternate at distal nodes; petiole 7–25 mm, strigose; blade usually narrowly to broadly elliptic, occasionally lance-elliptic, 10–100 × 5–35 mm, base cuneate to attenuate, margins coarsely crenate-dentate, strigose, revolute to nearly flat, apex broadly acute to acuminate, or obtuse, abaxial surface strigose with stiff, strongly tapered hairs, adaxial surface sparsely strigose-hirsute; venation pinnate, midvein prominent. |
Involucre | campanulate, 2.2–3 × 1.8–2 mm, glabrous; glands 4, crescent-shaped; 0.6–1 × 1–2 mm; horns slightly divergent to convergent, 0.1–0.2 mm. |
cylindric, 2.5–3 × 1.3–1.8 mm, glabrous; involucral lobes divided into 5–7 linear, papillate lobes; gland 1, yellow-green, sessile and broadly attached, 0.9 × 1.3 mm, opening oblong, glabrous; appendages absent. |
Staminate flowers | 15–20. |
5–8. |
Pistillate flowers | ovary glabrous; styles 1.2–2 mm, 2-fid. |
ovary glabrous or sparsely strigose; styles 1 mm, 2-fid 1/2 to nearly entire length. |
Capsules | globose, 2–2.8 × 2.2–2.7 mm, 3-lobed; cocci rounded, smooth except finely granulate toward abaxial line, glabrous; columella 2.1–2.7 mm. |
broadly ovoid, 2.9–3.3 × 4–4.8 mm, 3-lobed, glabrous; columella 2.2–2.7 mm. |
Seeds | gray or whitish, ovoid-oblong, 2–2.1 × 1.2–1.3 mm, smooth; caruncle ± rounded and flattened, 0.8 × 0.6 mm. |
black to brown or pale gray, ovoid to triangular-ovoid, angular in cross section, 2.4–2.9 × 2.2–2.9 mm, low-tuberculate, tubercles irregularly arranged or in faint, transverse row; caruncle 0.9–1.1 mm. |
Cyathial | arrangement: terminal pleiochasial branches 8–15, 1–2 times 2-branched; pleiochasial bracts similar in shape but shorter and narrower than distal leaves; dichasial bracts distinct, rhombic to reniform, base obtuse, margins entire, apex obtuse, mucronate; axillary cymose branches 12–23. |
arrangement: terminal pleiochasial branches usually 3, occasionally reduced to congested cyme, 1–2-branched; pleiochasial bracts 2–4, often whorled, green with diffuse greenish white to mauve near base, similar in shape and size to distal leaves or slightly narrower; dichasial bracts similar in shape to distal leaves but smaller, often highly reduced. |
Cyathia | peduncle 0–2 mm. |
peduncle 0.5–1 mm. |
2n | = 56. |
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Euphorbia agraria |
Euphorbia davidii |
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Phenology | Flowering and fruiting spring–summer. | Flowering and fruiting summer–fall. |
Habitat | Grasslands, roadside banks, pastures. | Forests, stream and riverbanks, prairies, roadsides and open disturbed areas. |
Elevation | 200–1600 m. (700–5200 ft.) | 200–1500 m. (700–4900 ft.) |
Distribution |
KS; MT; NE; NY; PA; WA; WY; AB; SK; Europe [Introduced in North America]
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AR; AZ; CA; CO; FL; IA; ID; IL; IN; KS; KY; LA; MA; MI; MN; MO; NC; NE; NJ; NM; NY; OH; OK; SD; TN; TX; UT; VA; VT; WI; WV; WY; ON; QC; Mexico (Chihuahua, Coahuila, Sonora) [Introduced in South America, Eurasia (China, Russia), Australia]
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Discussion | Euphorbia davidii is native from the southwestern United States and northern Mexico north through the southern Great Plains; it apparently is adventive elsewhere. The species is the weediest member of the E. dentata species group (following M. H. Mayfield 1997) and has become an agricultural weed in North America, South America (for example, Argentina), and in the Old World (particularly Australia and Russia). Euphorbia davidii can be distinguished from the closely similar E. dentata by its larger capsules and seeds, often more elliptic leaves, and shorter, stiffer hairs. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Source | FNA vol. 12, p. 298. | FNA vol. 12, p. 320. |
Parent taxa | Euphorbiaceae > Euphorbia > subg. Esula | Euphorbiaceae > Euphorbia > sect. Poinsettia |
Sibling taxa | ||
Synonyms | Tithymalus agrarius | |
Name authority | M. Bieberstein: Fl. Taur.-Caucas. 1: 375. (1808) | Subils: Kurtziana 17: 125, figs. 1, 2H–J. (1984) |
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