Euphorbia agraria |
Euphorbia carunculata |
|
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urban spurge |
sand-dune sandmat or spurge |
|
Habit | Herbs, perennial, with slender, spreading rootstock. | Herbs, annual, with taproot. |
Stems | erect or ascending, unbranched or branched, 30–90 cm, glabrous. |
prostrate, spreading and lanky or occasionally mat-forming, ± succulent, 70–150 cm, glabrous. |
Leaves | petiole absent; blade oblong-elliptic, 20–65 × 9–20 mm, base truncate to auriculate, margins entire, apex obtuse to rounded, surfaces glabrous; venation conspicuously pinnate, midvein prominent. |
opposite; stipules usually distinct, occasionally connate basally (primarily at distal nodes), usually divided into 2–5 subulate to subulate-filiform segments, occasionally forming narrow deltate segments (primarily at distal nodes), 0.8–1.8 mm, glabrous; petiole 3.1–6.3 mm, glabrous; blade ovate to elliptic-oblong, 5–26 × 4–12 mm, base subsymmetric to symmetric, rounded to cuneate, margins entire, apex usually mucronate, rarely acute or obtuse, surfaces glabrous; pinnately veined. |
Involucre | campanulate, 2.2–3 × 1.8–2 mm, glabrous; glands 4, crescent-shaped; 0.6–1 × 1–2 mm; horns slightly divergent to convergent, 0.1–0.2 mm. |
campanulate, 1.1–1.8 × 1.5–2.5 mm, glabrous; glands 4, yellowish, sessile or short stipitate, circular to oblong, 0.5–0.7 × 0.5–0.8 mm; appendages white to yellowish, ovate to oblong, occasionally rudimentary, (0–)0.8–1.2 × 0.8–1.5 mm, distal margin entire. |
Staminate flowers | 15–20. |
15–25. |
Pistillate flowers | ovary glabrous; styles 1.2–2 mm, 2-fid. |
ovary glabrous; styles 0.7–1 mm, 2-fid 1/2 length. |
Capsules | globose, 2–2.8 × 2.2–2.7 mm, 3-lobed; cocci rounded, smooth except finely granulate toward abaxial line, glabrous; columella 2.1–2.7 mm. |
ovoid, 4.7–5.5(–6) × 3.6–5.1 mm, glabrous; columella 4.3–5.1 mm. |
Seeds | gray or whitish, ovoid-oblong, 2–2.1 × 1.2–1.3 mm, smooth; caruncle ± rounded and flattened, 0.8 × 0.6 mm. |
grayish white to reddish brown mottled, bottle-shaped, strongly dorsiventrally compressed and weakly 3-angled in cross section, (2.8–)4.1–5.2 × 1.2–2(–3.4) mm, smooth; carunclelike structure linear, 0.4–0.5 × 0.1–0.2 mm. |
Cyathial | arrangement: terminal pleiochasial branches 8–15, 1–2 times 2-branched; pleiochasial bracts similar in shape but shorter and narrower than distal leaves; dichasial bracts distinct, rhombic to reniform, base obtuse, margins entire, apex obtuse, mucronate; axillary cymose branches 12–23. |
|
Cyathia | peduncle 0–2 mm. |
solitary at distal nodes; peduncle 1.4–3.6 mm. |
Euphorbia agraria |
Euphorbia carunculata |
|
Phenology | Flowering and fruiting spring–summer. | Flowering and fruiting summer–fall. |
Habitat | Grasslands, roadside banks, pastures. | Sand dunes. |
Elevation | 200–1600 m. (700–5200 ft.) | 400–1300 m. (1300–4300 ft.) |
Distribution |
KS; MT; NE; NY; PA; WA; WY; AB; SK; Europe [Introduced in North America]
|
KS; NM; OK; TX; Mexico (Chihuahua) |
Discussion | Euphorbia carunculata has a highly localized and scattered distribution. The species is restricted to sand dunes and known from only a handful of localities throughout its relatively wide range. The seeds are unique in being bottle-shaped and strongly laterally compressed, and unlike other members of sect. Anisophyllum, there is a minute, linear, carunclelike protuberance at the hilum. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
|
Source | FNA vol. 12, p. 298. | FNA vol. 12, p. 262. |
Parent taxa | Euphorbiaceae > Euphorbia > subg. Esula | Euphorbiaceae > Euphorbia > sect. Anisophyllum |
Sibling taxa | ||
Synonyms | Tithymalus agrarius | Chamaesyce carunculata |
Name authority | M. Bieberstein: Fl. Taur.-Caucas. 1: 375. (1808) | Waterfall: Rhodora 50: 63. (1948) |
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