Euphorbia agraria |
Euphorbia angusta |
|
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urban spurge |
Blackfoot sandmat, narrow-leaf spurge |
|
Habit | Herbs, perennial, with slender, spreading rootstock. | Herbs, perennial, with moderately to strongly thickened rootstock. |
Stems | erect or ascending, unbranched or branched, 30–90 cm, glabrous. |
erect, 12–43 cm, uniformly strigose. |
Leaves | petiole absent; blade oblong-elliptic, 20–65 × 9–20 mm, base truncate to auriculate, margins entire, apex obtuse to rounded, surfaces glabrous; venation conspicuously pinnate, midvein prominent. |
opposite; stipules distinct, linear-subulate or nodiform to papilliform nodiform to papilliform stipules often reddish brown, 0.1–0.7 mm, strigose; petiole 0.3–1.2 mm, strigose; blade: proximal ovate to ovate-elliptic, distal linear to elliptic-linear, 7–41 × 2–5 mm, distal leaf blades more than 6 times as long as wide, base asymmetric, cuneate to rounded, margins entire, often involute on drying, apex acute, surfaces usually short strigose, occasionally glabrous adaxially; venation pinnate, only midvein conspicuous. |
Involucre | campanulate, 2.2–3 × 1.8–2 mm, glabrous; glands 4, crescent-shaped; 0.6–1 × 1–2 mm; horns slightly divergent to convergent, 0.1–0.2 mm. |
turbinate to campanulate-turbinate, 1–1.5 × 1–1.4 mm, strigose; glands 4, green to yellow-green, concave, narrowly oblong, 0.2–0.4 × 0.4–0.7 mm; appendages white, flabellate, 0.5–1.1 × 0.3–0.5 mm, distal margin shallowly and irregularly toothed. |
Staminate flowers | 15–20. |
16–26. |
Pistillate flowers | ovary glabrous; styles 1.2–2 mm, 2-fid. |
ovary strigose; styles 0.3–0.5 mm, 2-fid at apex to almost 1/2 length. |
Capsules | globose, 2–2.8 × 2.2–2.7 mm, 3-lobed; cocci rounded, smooth except finely granulate toward abaxial line, glabrous; columella 2.1–2.7 mm. |
broadly ovoid, 2–2.6 × 2.5–3 mm, strigose; columella 1.6–2.2 mm. |
Seeds | gray or whitish, ovoid-oblong, 2–2.1 × 1.2–1.3 mm, smooth; caruncle ± rounded and flattened, 0.8 × 0.6 mm. |
white, ovoid, 4-angled in cross section, 1.7–2.2 × 1.1–1.2 mm, transversely low-ridged or wrinkled. |
Cyathial | arrangement: terminal pleiochasial branches 8–15, 1–2 times 2-branched; pleiochasial bracts similar in shape but shorter and narrower than distal leaves; dichasial bracts distinct, rhombic to reniform, base obtuse, margins entire, apex obtuse, mucronate; axillary cymose branches 12–23. |
|
Cyathia | peduncle 0–2 mm. |
solitary at distal nodes; peduncle 1.1–2.4 mm. |
2n | = 28. |
|
Euphorbia agraria |
Euphorbia angusta |
|
Phenology | Flowering and fruiting spring–summer. | Flowering and fruiting early spring–fall. |
Habitat | Grasslands, roadside banks, pastures. | Rocky limestone soils. |
Elevation | 200–1600 m. (700–5200 ft.) | 400–1200 m. (1300–3900 ft.) |
Distribution |
KS; MT; NE; NY; PA; WA; WY; AB; SK; Europe [Introduced in North America]
|
TX; Mexico (Coahuila) |
Discussion | Euphorbia angusta, which in the flora area is known from the trans-Pecos region to the Edwards Plateau, is easily recognized by its erect habit, linear leaves, and relatively showy involucral gland appendages with toothed margins. The species is closely related to E. acuta and the Mexican endemic E. johnstonii Mayfield (M. H. Mayfield 1991); it is not only morphologically distinctive but is also the only species in sect. Anisophyllum with C3 photosynthesis (R. F. Sage et al. 2011; T. L. Sage et al. 2011; G. L. Webster 1975). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
|
Source | FNA vol. 12, p. 298. | FNA vol. 12, p. 260. |
Parent taxa | Euphorbiaceae > Euphorbia > subg. Esula | Euphorbiaceae > Euphorbia > sect. Anisophyllum |
Sibling taxa | ||
Synonyms | Tithymalus agrarius | Chamaesyce angusta |
Name authority | M. Bieberstein: Fl. Taur.-Caucas. 1: 375. (1808) | Engelmann: in W. H. Emory, Rep. U.S. Mex. Bound. 2(1): 189. (1859) |
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