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Abrams' sandmat, Abrams' spurge

rocklands spurge, Southern Florida sandmat

Habit Herbs, annual, with slender taproot. Herbs, perennial, with moderately to strongly thickened rootstock.
Stems

prostrate, mat-forming, 10–35(–50) cm, shortly pilose or puberulent at least proximally, often glabrous distally.

prostrate to ascending, occasionally mat-forming, terete to slightly flattened, 5–18 cm, glabrous on lower surface, strigillose to sericeous on upper surface.

Leaves

opposite;

stipules distinct, divided into 5–7 subulate-filiform segments, 0.6–1.1 mm, usually glabrous, rarely pilose;

petiole 0.5–1 mm, glabrous;

blade ovate, elliptic-oblong, or slightly ovate-cordate, 3–11 × 2–5 mm, base asymmetric, truncate to hemicordate, margins serrulate at least toward apex, often entire toward base, apex acute to obtuse, surfaces sometimes with red spot in center, glabrous; usually only the midvein conspicuous.

opposite;

stipules distinct, subulate, better developed on lower side of stem, 0.4–1.3 mm, glabrous or pilose;

petiole 0.3–0.5 mm, puberulent, sericeous, or strigillose;

blade oblong to ovate, 4–7 × 2–4 mm, base asymmetric, hemicordate, larger side sometimes amplexicaulous, margins entire or serrulate, apex obtuse, surfaces puberulent, sericeous, or strigillose; 3-veined from base, lateral veins inconspicuous.

Involucre

obconic, 0.5–0.6 × 0.4–0.5 mm, glabrous;

glands 4, yellowish to pink, circular to oblong, 0.1 × 0.1–0.2 mm;

appendages absent, or white to pink, semicircular to broadly ovate, to 0.1 × 0.2 mm, distal margin entire or shallowly lobed.

campanulate, 1–1.3 × 1–1.1 mm, canescent;

glands 4, pinkish, oblong or reniform, 0.2 × 0.4–0.6 mm;

appendages white to pink, lunate, unequal, those of proximal glands oblique, 0.6–1 × 1.4–1.6 mm, those of distal glands symmetric, 0.3 × 0.8–0.9 mm, irregularly undulate to incised.

Staminate flowers

3–5.

15.

Pistillate flowers

ovary glabrous;

styles 0.1–0.3 mm, 2-fid nearly entire length.

ovary puberulent, sericeous or strigillose;

styles 0.5–1 mm, 2-fid at apex.

Capsules

ellipsoid to ovoid, 1.3–1.5 × 1.1–1.5 mm, glabrous;

columella 1–1.3 mm.

oblate, well exserted from involucre at maturity, 1–1.3 × 1.3–1.8 mm, puberulent, sericeous or strigillose;

columella 1 mm.

Seeds

light gray to light brown, narrowly ovoid to ovoid, 4-angled in cross section, 1–1.2 × 0.6–0.7 mm, with 3–5 prominent transverse ridges that often interrupt abaxial keel.

whitish to gray, ovoid, 4-angled in cross section, 0.7–0.8 × 0.5–0.6 mm, with 3–4 transverse sulci alternating with low transverse ridges.

Cyathia

solitary at distal nodes of primary stems or at nodes of short congested axillary branchlets;

peduncle 0.2–0.5 mm.

solitary at distal nodes or at nodes of short, congested, axillary shoots;

peduncle to 0.5 mm.

Euphorbia abramsiana

Euphorbia pergamena

Phenology Flowering and fruiting summer–fall. Flowering and fruiting year-round.
Habitat Desert scrub and desert grasslands. Crevices of limestone outcrops in pinelands and pine-palm woods.
Elevation -40–1400 m. (-100–4600 ft.) 0–10 m. (0–0 ft.)
Distribution
from FNA
AZ; CA; NM; TX; Mexico (Baja California, Baja California Sur, Sonora)
[WildflowerSearch map]
[BONAP county map]
from FNA
FL; West Indies (Cuba, Hispaniola)
Discussion

Euphorbia pergamena is a Florida-listed threatened species known within the flora area from only Miami-Dade and Monroe counties in extreme southern Florida. The species is often included as a subspecies of E. adenoptera (for example, D. G. Burch 1965), but here A. Herndon (1993b) is followed and it is treated as a distinct species.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 12, p. 259. FNA vol. 12, p. 282.
Parent taxa Euphorbiaceae > Euphorbia > sect. Anisophyllum Euphorbiaceae > Euphorbia > sect. Anisophyllum
Sibling taxa
E. aaron-rossii, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana
E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana
Synonyms Chamaesyce abramsiana Chamaesyce adenoptera subsp. pergamena, C. pergamena, E. adenoptera subsp. pergamena
Name authority L. C. Wheeler: Bull. S. Calif. Acad. Sci. 33: 109. (1934) Small: Bull. Torrey Bot. Club 25: 615. (1898)
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