Euphorbia abramsiana |
Euphorbia bombensis |
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Abrams' sandmat, Abrams' spurge |
Dixie sandmat, southern seaside spurge |
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Habit | Herbs, annual, with slender taproot. | Herbs, usually annual, rarely perennial, with taproot. |
Stems | prostrate, mat-forming, 10–35(–50) cm, shortly pilose or puberulent at least proximally, often glabrous distally. |
prostrate or slightly ascending, 10–40 cm, glabrous. |
Leaves | opposite; stipules distinct, divided into 5–7 subulate-filiform segments, 0.6–1.1 mm, usually glabrous, rarely pilose; petiole 0.5–1 mm, glabrous; blade ovate, elliptic-oblong, or slightly ovate-cordate, 3–11 × 2–5 mm, base asymmetric, truncate to hemicordate, margins serrulate at least toward apex, often entire toward base, apex acute to obtuse, surfaces sometimes with red spot in center, glabrous; usually only the midvein conspicuous. |
opposite; stipules distinct, linear-subulate, usually divided into 3 linear segments, 1–2 mm, glabrous; petiole 1–2 mm, glabrous; blade oblong or elliptic-oblong, 4–15 × 2–3 mm, base asymmetric to nearly symmetric, obtuse, margins entire, apex acute to mucronate, surfaces green to reddish flushed, glabrous; only midvein conspicuous. |
Involucre | obconic, 0.5–0.6 × 0.4–0.5 mm, glabrous; glands 4, yellowish to pink, circular to oblong, 0.1 × 0.1–0.2 mm; appendages absent, or white to pink, semicircular to broadly ovate, to 0.1 × 0.2 mm, distal margin entire or shallowly lobed. |
obconic-campanulate, 1.2–1.6 × 1.5–1.7 mm, glabrous; glands 4, green to red, slightly concave, elliptic, oblong, or subcircular, 0.3–0.5 × 0.4–0.6 mm; appendages white or pink, semilunate, fringing edge of gland, sometimes rudimentary, (0–)0.1–0.5(–0.7) × 0.5–0.8 mm, distal margin crenate to entire. |
Staminate flowers | 3–5. |
5–16. |
Pistillate flowers | ovary glabrous; styles 0.1–0.3 mm, 2-fid nearly entire length. |
ovary glabrous; styles 0.2–0.3 mm, 2-fid 1/2 length. |
Capsules | ellipsoid to ovoid, 1.3–1.5 × 1.1–1.5 mm, glabrous; columella 1–1.3 mm. |
broadly ovoid, 2–2.1 × 2.3–2.5 mm, glabrous; columella 1.5–2 mm. |
Seeds | light gray to light brown, narrowly ovoid to ovoid, 4-angled in cross section, 1–1.2 × 0.6–0.7 mm, with 3–5 prominent transverse ridges that often interrupt abaxial keel. |
ashy white, plumply ovoid, terete to bluntly subangled in cross section, 1.5–1.9 × 1–1.2 mm, smooth or minutely pitted, with smooth brown line from top to bottom on adaxial side. |
Cyathia | solitary at distal nodes of primary stems or at nodes of short congested axillary branchlets; peduncle 0.2–0.5 mm. |
solitary or in small, cymose clusters at distal nodes; peduncle 0.5–3 mm. |
Euphorbia abramsiana |
Euphorbia bombensis |
|
Phenology | Flowering and fruiting summer–fall. | Flowering and fruiting spring–fall. |
Habitat | Desert scrub and desert grasslands. | Coastal dunes and sandy habitats. |
Elevation | -40–1400 m. (-100–4600 ft.) | 0–30 m. (0–100 ft.) |
Distribution |
AZ; CA; NM; TX; Mexico (Baja California, Baja California Sur, Sonora)
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AL; FL; GA; LA; MS; NC; SC; TX; VA; Mexico; Central America; n South America (Venezuela)
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Discussion | Euphorbia bombensis is similar and closely related to E. cumulicola, E. geyeri, and E. polygonifolia. It differs notably from E. polygonifolia in its smaller, plumply ovoid seeds. L. C. Wheeler (1941) suggested that where E. bombensis and E. polygonifolia are sympatric, E. bombensis grows farther away from the shore. Euphorbia bombensis is usually distinguished from E. cumulicola by the latter's smaller, isomorphic leaves that lack any fleshiness, smaller seeds, and diffuse growth habit. Euphorbia bombensis differs from E. geyeri in its usually shorter, less conspicuous involucral gland appendages and its geographic restriction to the coastal plain. However, Wheeler pointed out that plants of E. bombensis from Texas have more or less conspicuous involucral gland appendages. Examination of specimens confirmed that E. bombensis occasionally has conspicuous involucral gland appendages, and because of this, E. bombensis and E. geyeri are difficult to distinguish in Texas. This clade of closely related, sand and dune specialists requires further study. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Source | FNA vol. 12, p. 259. | FNA vol. 12, p. 261. |
Parent taxa | Euphorbiaceae > Euphorbia > sect. Anisophyllum | Euphorbiaceae > Euphorbia > sect. Anisophyllum |
Sibling taxa | ||
Synonyms | Chamaesyce abramsiana | Chamaesyce ammannioides, C. bombensis, C. ingallsii, E. ammannioides |
Name authority | L. C. Wheeler: Bull. S. Calif. Acad. Sci. 33: 109. (1934) | Jacquin: Enum. Syst. Pl., 22. (1760) |
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