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round-leaf boneset, round-leaf thoroughwort, roundleaf eupatorium

white thoroughwort

Habit Perennials, 40–100+ cm. Perennials, 40–100+ cm.
Stems

(from short rhizomes) single, sparsely branched distally, puberulent.

(from short caudices or stout rhizomes) single, sparsely branched distally, pubescent throughout.

Leaves

usually opposite (distal sometimes alternate, lateral buds dormant or producing 1 pair of leaves);

sessile or subsessile;

blades ± 3-nerved, usually ± deltate to orbiculate, sometimes ovate, 15–50(–70) × 15–40(–60) mm (lengths mostly 1–2 times widths), bases broadly rounded to truncate, margins crenate to serrate, apices acute, faces puberulent to villous, gland-dotted.

usually opposite (distal sometimes alternate);

sessile;

blades ± 3-nerved or pinnately nerved, elliptic to oblanceolate, 30–120 × 10–40 mm (lengths mostly 3–4 times widths), bases narrowly cuneate (sometimes oblique), margins subentire, serrate, or serrulate, apices rounded to acute, faces setulose (denser on midribs and veinlets) to glabrate, usually gland-dotted.

Florets

5;

corollas 3–4 mm.

(4–)5;

corollas 4–4.5 mm.

Phyllaries

8–10 in 2–3 series, elliptic to oblanceolate, 2–6 × 0.5–1.5 mm, apices acute, abaxial faces puberulent, gland-dotted.

9–15 in 2–4 series, linear, 1.5–9 × 0.6–1 mm, apices acuminate to attenuate, sometimes mucronate, abaxial faces pubescent throughout, gland-dotted.

Heads

in corymbiform arrays.

in corymbiform arrays.

Cypselae

2–3 mm;

pappi of 30–40 bristles 3.5–4 mm.

2.5–3.5 mm;

pappi of 40–50 bristles 3.5–4.5 mm.

Eupatorium rotundifolium

Eupatorium album

Distribution
from FNA
AL; AR; CT; DC; DE; FL; GA; IN; KY; LA; MA; MD; MO; MS; NC; NH; NJ; NY; OH; OK; PA; RI; SC; TN; TX; VA; WV
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[BONAP county map]
from FNA
AL; CT; DC; DE; FL; GA; KY; LA; MD; MS; NC; NJ; NY; OH; PA; SC; TN; VA; WV
[WildflowerSearch map]
[BONAP county map]
Discussion

Varieties 3 (3 in the flora).

Eupatorium rotundifolium is one of the more variable species of the genus; its apparent propensity to hybridize with other species has led it to be described as a compilospecies and has complicated its delimitation. Hybrids with E. perfoliatum and E. sessilifolium are sufficiently distinctive to be treated here as distinctive entities (E. ×cordigerum and E. godfreyanum, respectively), and the varieties ovatum and scabridum have also been proposed to be of hybrid origin. The boundaries between varieties are indistinct and individual specimens may be difficult to place with confidence.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 3 (3 in the flora).

Eupatorium album is represented by widespread diploid populations and by polyploid, apomictic populations, segregated as varieties, that may have arisen by hybridization with other species. Molecular data suggest that var. vaseyi arose via hybridization between var. album and E. sessilifolium. They also suggest that E. album var. subvenosum arose from hybridization between E. album var. album and E. serotinum.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Leaf blades 3-nerved from bases, bases subtruncate to broadly cuneate, margins crenate
var. rotundifolium
1. Leaf blades 3-nerved distal to bases, bases broadly cuneate to cuneate, margins serrate
→ 2
2. Leaf blades 30–70 × 30–60 mm, broadest nearmiddles
var. ovatum
2. Leaf blades 20–50 × 15–30 mm, broadest nearbases
var. scabridum
1. Leaf blades: margins coarsely serrate, apices obtuse to rounded, faces pubescent
var. album
1. Leaf blades: margins subentire, serrulate, or serrate, apices acute, faces pubescent to glabrate
→ 2
2. Leaf blades 3-nerved from bases, 40–70 × 10–20 mm, margins subentire to serrulate
var. subvenosum
2. Leaf blades 3-nerved distal to bases, 50–110 × 20–40 mm, margins evenly serrate
var. vaseyi
Source FNA vol. 21, p. 472. FNA vol. 21, p. 465.
Parent taxa Asteraceae > tribe Eupatorieae > Eupatorium Asteraceae > tribe Eupatorieae > Eupatorium
Sibling taxa
E. album, E. altissimum, E. anomalum, E. cannabinum, E. capillifolium, E. compositifolium, E. godfreyanum, E. hyssopifolium, E. lancifolium, E. leptophyllum, E. leucolepis, E. linearifolium, E. mikanioides, E. mohrii, E. perfoliatum, E. petaloideum, E. pilosum, E. resinosum, E. semiserratum, E. serotinum, E. sessilifolium, E. ×cordigerum, E. ×pinnatifidum
E. altissimum, E. anomalum, E. cannabinum, E. capillifolium, E. compositifolium, E. godfreyanum, E. hyssopifolium, E. lancifolium, E. leptophyllum, E. leucolepis, E. linearifolium, E. mikanioides, E. mohrii, E. perfoliatum, E. petaloideum, E. pilosum, E. resinosum, E. rotundifolium, E. semiserratum, E. serotinum, E. sessilifolium, E. ×cordigerum, E. ×pinnatifidum
Subordinate taxa
E. rotundifolium var. ovatum, E. rotundifolium var. rotundifolium, E. rotundifolium var. scabridum
E. album var. album, E. album var. subvenosum, E. album var. vaseyi
Name authority Linnaeus: Sp. Pl. 2: 837. (1753) Linnaeus: Mant. Pl., 111. (1767): Syst. Nat. ed. 12, 2: 536. (1767)
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