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hyssop-leaf boneset, hyssop-leaf thoroughwort

Mohr's thoroughwort

Habit Perennials, 50–100+ cm. Perennials, 30–100+ cm.
Stems

(from short caudices or rhizomes) single, sparsely branched distally, pubescent throughout.

(from tuberous rhizomes) multiple, densely branched distally, puberulent throughout.

Leaves

usually opposite or whorled (distal sometimes alternate, spreading or horizontal); simple, sessile;

blades 3-nerved from bases (laterals sometimes weak), lance-linear, lance-oblong, or linear, 20–60 × 2–15 mm (lengths mostly 6–40 times widths), bases cuneate, margins entire, laciniate-serrate, or serrate, apices acute, faces scabrous (at least abaxial).

usually opposite (distal sometimes alternate);

sessile or subsessile;

blades 3-nerved distal to bases, oblanceolate, 20–80 × 5–10(–20) mm, bases cuneate, margins serrate proximally, entire distally, apices acute, faces puberulent (abaxial), glabrous or glabrate (adaxial), gland-dotted.

Florets

5;

corollas 3–3.5 mm.

5;

corollas 2–4 mm.

Phyllaries

8–10 in 2–3 series, elliptic to oblong, 1.5–5 × 1–1.5 mm, apices obtuse to acute (not mucronate), abaxial faces pubescent throughout (more densely distally).

7–10 in 1–2 series, oblanceolate, 1–3 × 0.2–0.5 mm, apices rounded, abaxial faces puberulent, gland-dotted.

Heads

in corymbiform arrays.

in corymbiform arrays.

Cypselae

2–3 mm;

pappi of 20–30 bristles 3.5–4 mm.

1–2 mm;

pappi of 20–30 bristles 2.5–3 mm.

2n

= 20, 30, 40.

Eupatorium hyssopifolium

Eupatorium mohrii

Phenology Flowering Jul–Sep.
Habitat Moist, low ground, margins of ponds, sandy soils
Elevation 10–100+ m (0–300+ ft)
Distribution
from FNA
AL; AR; CT; DE; FL; GA; IL; KY; LA; MA; MD; MO; MS; NC; NJ; NY; OH; PA; RI; SC; TN; TX; VA; WI; WV
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; FL; GA; LA; MS; NC; SC; TX; VA
[WildflowerSearch map]
[BONAP county map]
Discussion

Varieties 2 (2 in the flora).

Eupatorium hyssopifolium includes both diploid and polyploid cytotypes and presents a complex situation taxonomically. The diploids are placed here in var. hyssopifolium, which also includes polyploids and is characterized by relatively narrow leaves. Variety laciniatum has broader leaves and molecular data suggest that it arose through hybridization between var. hyssopifolium and E. serotinum; the name E. torreyanum has been applied to plants of similar morphology, but these are hybrid derivatives of E. serotinum and E. mohrii. It has also been suggested that var. hyssopifolium has hybridized with E. album var. album to form E. saltuense, and with diploids of E. linearifolium to form polyploid populations included here under E. linearifolium. Eupatorium hyssopifolium is characterized by a tendency for the leaves to be whorled, in 3s and 4s, and for axillary buds to produce leaves without expansion of shoots; the combination of whorled leaves and suppressed axillary shoots gives the nodes a somewhat tufted or verticillate appearance.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

As treated here, Eupatorium mohrii includes both sexual diploid (sometimes segregated as E. recurvans) and apomictic polyploid populations that are suggested by molecular data to be autoploids. Eupatorium saltuense refers to hybrids shown by molecular data to involve E. mohrii and E. serotinum. Molecular data also document frequent hybridization with E. rotundifolium.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Leaves mostly linear, 2–5 mm wide, margins usually entire or obscurely serrulate
var. hyssopifolium
1. Leaves mostly lance-linear to lanceolate, 5–15 mm wide, margins usually laciniately toothed
var. laciniatum
Source FNA vol. 21, p. 468. FNA vol. 21, p. 470.
Parent taxa Asteraceae > tribe Eupatorieae > Eupatorium Asteraceae > tribe Eupatorieae > Eupatorium
Sibling taxa
E. album, E. altissimum, E. anomalum, E. cannabinum, E. capillifolium, E. compositifolium, E. godfreyanum, E. lancifolium, E. leptophyllum, E. leucolepis, E. linearifolium, E. mikanioides, E. mohrii, E. perfoliatum, E. petaloideum, E. pilosum, E. resinosum, E. rotundifolium, E. semiserratum, E. serotinum, E. sessilifolium, E. ×cordigerum, E. ×pinnatifidum
E. album, E. altissimum, E. anomalum, E. cannabinum, E. capillifolium, E. compositifolium, E. godfreyanum, E. hyssopifolium, E. lancifolium, E. leptophyllum, E. leucolepis, E. linearifolium, E. mikanioides, E. perfoliatum, E. petaloideum, E. pilosum, E. resinosum, E. rotundifolium, E. semiserratum, E. serotinum, E. sessilifolium, E. ×cordigerum, E. ×pinnatifidum
Subordinate taxa
E. hyssopifolium var. hyssopifolium, E. hyssopifolium var. laciniatum
Synonyms E. recurvans
Name authority Linnaeus: Sp. Pl. 2: 836. (1753) Greene: Contr. U.S. Natl. Herb. 6: 762, plate 11. (1901)
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