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hyssop-leaf boneset, hyssop-leaf thoroughwort

false fennel

Habit Perennials, 50–100+ cm. Perennials, 50–200 cm.
Stems

(from short caudices or rhizomes) single, sparsely branched distally, pubescent throughout.

(from short caudices) single, branched distally, glabrous throughout, gland-dotted.

Leaves

usually opposite or whorled (distal sometimes alternate, spreading or horizontal); simple, sessile;

blades 3-nerved from bases (laterals sometimes weak), lance-linear, lance-oblong, or linear, 20–60 × 2–15 mm (lengths mostly 6–40 times widths), bases cuneate, margins entire, laciniate-serrate, or serrate, apices acute, faces scabrous (at least abaxial).

opposite (proximal) or alternate (nodes often appearing leafy by development of leaves on lateral buds without axis elongation); petiolate or sessile;

blades (usually pinnately or ternately lobed) or lobes 1-nerved or pinnately nerved, linear, 20–100 × 0.2–0.5(–1) mm, bases slightly expanded, margins entire, apices rounded to acute, faces sparsely puberulent to glabrate, gland-dotted.

Florets

5;

corollas 3–3.5 mm.

5;

corollas 2–2.5 mm.

Phyllaries

8–10 in 2–3 series, elliptic to oblong, 1.5–5 × 1–1.5 mm, apices obtuse to acute (not mucronate), abaxial faces pubescent throughout (more densely distally).

6–10 in 1–2 series, lanceolate, 1.5–3 × 0.5–0.7 mm, apices acuminate, mucronate, abaxial faces glabrous or glabrate (not gland-dotted).

Heads

in corymbiform arrays.

in paniculiform arrays (branches supporting heads recurved, secund).

Cypselae

2–3 mm;

pappi of 20–30 bristles 3.5–4 mm.

1–1.5 mm;

pappi of 15–30 bristles 2–2.5 mm.

2n

= 20.

Eupatorium hyssopifolium

Eupatorium leptophyllum

Phenology Flowering Aug–Oct.
Habitat Pond margins, wet, low places, shallow water
Elevation 10–50+ m (0–200+ ft)
Distribution
from FNA
AL; AR; CT; DE; FL; GA; IL; KY; LA; MA; MD; MO; MS; NC; NJ; NY; OH; PA; RI; SC; TN; TX; VA; WI; WV
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; FL; GA; MS; NC; SC
[WildflowerSearch map]
[BONAP county map]
Discussion

Varieties 2 (2 in the flora).

Eupatorium hyssopifolium includes both diploid and polyploid cytotypes and presents a complex situation taxonomically. The diploids are placed here in var. hyssopifolium, which also includes polyploids and is characterized by relatively narrow leaves. Variety laciniatum has broader leaves and molecular data suggest that it arose through hybridization between var. hyssopifolium and E. serotinum; the name E. torreyanum has been applied to plants of similar morphology, but these are hybrid derivatives of E. serotinum and E. mohrii. It has also been suggested that var. hyssopifolium has hybridized with E. album var. album to form E. saltuense, and with diploids of E. linearifolium to form polyploid populations included here under E. linearifolium. Eupatorium hyssopifolium is characterized by a tendency for the leaves to be whorled, in 3s and 4s, and for axillary buds to produce leaves without expansion of shoots; the combination of whorled leaves and suppressed axillary shoots gives the nodes a somewhat tufted or verticillate appearance.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Leaves mostly linear, 2–5 mm wide, margins usually entire or obscurely serrulate
var. hyssopifolium
1. Leaves mostly lance-linear to lanceolate, 5–15 mm wide, margins usually laciniately toothed
var. laciniatum
Source FNA vol. 21, p. 468. FNA vol. 21, p. 469.
Parent taxa Asteraceae > tribe Eupatorieae > Eupatorium Asteraceae > tribe Eupatorieae > Eupatorium
Sibling taxa
E. album, E. altissimum, E. anomalum, E. cannabinum, E. capillifolium, E. compositifolium, E. godfreyanum, E. lancifolium, E. leptophyllum, E. leucolepis, E. linearifolium, E. mikanioides, E. mohrii, E. perfoliatum, E. petaloideum, E. pilosum, E. resinosum, E. rotundifolium, E. semiserratum, E. serotinum, E. sessilifolium, E. ×cordigerum, E. ×pinnatifidum
E. album, E. altissimum, E. anomalum, E. cannabinum, E. capillifolium, E. compositifolium, E. godfreyanum, E. hyssopifolium, E. lancifolium, E. leucolepis, E. linearifolium, E. mikanioides, E. mohrii, E. perfoliatum, E. petaloideum, E. pilosum, E. resinosum, E. rotundifolium, E. semiserratum, E. serotinum, E. sessilifolium, E. ×cordigerum, E. ×pinnatifidum
Subordinate taxa
E. hyssopifolium var. hyssopifolium, E. hyssopifolium var. laciniatum
Synonyms E. capillifolium var. leptophyllum
Name authority Linnaeus: Sp. Pl. 2: 836. (1753) de Candolle: in A. P. de Candolle and A. L. P. P. de Candolle, Prodr. 5: 176. (1836)
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