Eulobus
Onagraceae subfam. onagroideae
erect [to prostrate], unbranched or branched.
cauline, sometimes also in basal rosette, alternate;
stipules absent;
sessile or petiolate;
blade pinnatifid to lobed, [subentire].
stipules present or absent.
open spikes, erect.
bisexual, actinomorphic, buds erect;
floral tube relatively short, deciduous (with sepals, petals, and stamens) after anthesis, lined with a lobed, fleshy nectary disc [or with rounded, fleshy disc at base of style];
sepals 4, reflexed separately, in pairs, or as a unit, (rarely spreading);
petals 4, yellow, fading orangish red, usually finely flecked with red near base;
stamens 8, in 2 subequal series or epipetalous series shorter, anthers versatile, pollen shed singly;
ovary 4-locular, stigma globose to cylindrical, surface wet and non-papillate.
floral tube present or, rarely, absent;
sepals 2 or 4 (very rarely 3), deciduous with floral tube, petals, and stamens;
petals yellow, white, pink, red, rarely in combination.
a capsule, straight or slightly curved [contorted], somewhat torulose or subterete, loculicidally dehiscent, midrib of each valve prominent;
sessile.
numerous, in 1 row per locule, narrowly obovoid, ± triangular in cross section, finely papillose.
Eulobus
Onagraceae subfam. onagroideae
Species 4 (1 in the flora).
Eulobus is characterized by olive brown seeds with distinct purple spots, yellow petals with maroon flecks near the base, the distal part ultraviolet-reflective, leaves mostly lobed or pinnatifid, and capsules somewhat contorted or straight, and often sharply reflexed at maturity. R. A. Levin et al. (2004) included E. californicus and E. crassifolius (Greene) W. L. Wagner & Hoch (a species of Baja California, Mexico) in their molecular analysis and found Eulobus to be strongly supported as monophyletic; they also found that Eulobus plus (Chylismia plus Oenothera) formed a weakly supported clade. Reproductive features include: self-incompatible [E. crassifolius, E. sceptrostigma (Brandegee) W. L. Wagner & Hoch] or self-compatible [E. angelorum (S. Watson) W. L. Wagner & Hoch, E. californicus], flowers diurnal, outcrossing and pollinated mainly by small oligolectic bees (E. G. Linsley 1963b, 1973) or autogamous (P. H. Raven 1969).
(Discussion copyrighted by Flora of North America; reprinted with permission.)
Genera 21, species 582 (16 genera, 246 species in the flora).
Onagroideae encompass the main lineage of the family, after the early branching of Ludwigia (R. A. Levin et al. 2003, 2004). This large and diverse lineage is distinguished by the presence of a floral tube beyond the apex of the ovary; sepals deciduous with the floral tube, petals, and stamens; pollen shed in monads (or tetrads in Chylismia sect. Lignothera and all but one species of Epilobium); ovular vascular system exclusively transseptal (R. H. Eyde 1981); ovule archesporium multicellular (H. Tobe and P. H. Raven 1996); and change in base chromosome number from x = 8 in Ludwigia to x = 10 or x = 11 at the base of Onagroideae (Raven 1979; Levin et al. 2003). Molecular work (Levin et al. 2003, 2004) substantially supports the traditional tribal classification (P. A. Munz 1965; Raven 1979, 1988); tribes are recognized to delimit major branches within the phylogeny of Onagroideae, where the branches comprise strongly supported monophyletic groups of one or more genera.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
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