Flora of North America species comparison

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Eulobus
Habit Herbs, annual, [perennial, subshrubs], caulescent; with a taproot.
Stems

erect [to prostrate], unbranched or branched.
Leaves

cauline, sometimes also in basal rosette, alternate;

stipules absent;

sessile or petiolate;

blade pinnatifid to lobed, [subentire].
Inflorescences

open spikes, erect.
Flowers

bisexual, actinomorphic, buds erect;

floral tube relatively short, deciduous (with sepals, petals, and stamens) after anthesis, lined with a lobed, fleshy nectary disc [or with rounded, fleshy disc at base of style];

sepals 4, reflexed separately, in pairs, or as a unit, (rarely spreading);

petals 4, yellow, fading orangish red, usually finely flecked with red near base;

stamens 8, in 2 subequal series or epipetalous series shorter, anthers versatile, pollen shed singly;

ovary 4-locular, stigma globose to cylindrical, surface wet and non-papillate.
Fruit

a capsule, straight or slightly curved [contorted], somewhat torulose or subterete, loculicidally dehiscent, midrib of each valve prominent;

sessile.
Seeds

numerous, in 1 row per locule, narrowly obovoid, ± triangular in cross section, finely papillose.
xI> = 7.

Eulobus
Distribution
sw United States; nw Mexico
[BONAP county map]
Discussion

Species 4 (1 in the flora).

Eulobus is characterized by olive brown seeds with distinct purple spots, yellow petals with maroon flecks near the base, the distal part ultraviolet-reflective, leaves mostly lobed or pinnatifid, and capsules somewhat contorted or straight, and often sharply reflexed at maturity. R. A. Levin et al. (2004) included E. californicus and E. crassifolius (Greene) W. L. Wagner & Hoch (a species of Baja California, Mexico) in their molecular analysis and found Eulobus to be strongly supported as monophyletic; they also found that Eulobus plus (Chylismia plus Oenothera) formed a weakly supported clade. Reproductive features include: self-incompatible [E. crassifolius, E. sceptrostigma (Brandegee) W. L. Wagner & Hoch] or self-compatible [E. angelorum (S. Watson) W. L. Wagner & Hoch, E. californicus], flowers diurnal, outcrossing and pollinated mainly by small oligolectic bees (E. G. Linsley 1963b, 1973) or autogamous (P. H. Raven 1969).

(Discussion copyrighted by Flora of North America; reprinted with permission.)
Source FNA vol. 10. Author: Warren L. Wagner.
Parent taxa Onagraceae > subfam. Onagroideae > tribe Onagreae
Subordinate taxa
E. californicus
Synonyms Camissonia section eulobus, Oenothera section eulobus, Oenothera subg. eulobus
Name authority Nuttall ex Torrey & A. Gray: Fl. N. Amer. 1: 514. (1840)
Web links 
Unless otherwise noted, all data comes from the online version of the Flora of North America, who have made most of their content available via a Creative Commons license (CC BY 4.0).



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