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cottonleaf, cudweed, euchiton

globe cotton-leaf, star-cudweed, tropical creeping cudweed

Habit Annuals or perennials, 5–80 cm (usually fibrous-rooted, sometimes rhizomatous, usually stoloniferous). Annuals, 5–80 cm; taprooted; stolons absent.
Aerial stems

simple or branched from bases (sometimes branched from leaf axils), thinly and persistently white-tomentose.

Stems

usually 1, erect.

Leaves

basal and cauline (sometimes in rosettes); alternate;

petiolate or sessile;

blades oblanceolate, spatulate, lanceolate, or linear, bases cuneate or ampliate, margins entire (sometimes undulate and/or revolute), faces bicolor, abaxial usually silvery, tomentose, adaxial usually green, glabrate or glabrous.

basal and proximal cauline withering before flowering (blades 1-nerved, oblanceolate to spatulate);

cauline 8–12, blades linear, 2–4 cm × 1–2 mm (largest at midstem), bases not clasping, margins revolute, sometimes undulate, abaxial faces white, tomentose, adaxial faces green, glabrous.

Bracts

subtending heads 4–8, 10–30 mm, surpassing heads.

Involucres

narrowly campanulate to cylindric, 3–5 mm.

3.5–4 mm.

Receptacles

flat, smooth, epaleate.

Pistillate florets

16–26.

Peripheral (pistillate) florets

16–150 (more numerous than bisexual);

corollas purple or distally purplish.

Bisexual florets

1.

Phyllaries

in 3–4+ series, mostly stramineous to brownish, sometimes purplish to pinkish (hyaline, stereomes not glandular), unequal, chartaceous toward tips.

brownish to tawny, sometimes purple-tinged (shiny), elliptic-lanceolate, apices acute.

Heads

disciform, usually in terminal clusters (subtended by leafy bracts, sometimes with axillary clusters), rarely borne singly.

in globose clusters 10–20 mm diam.

Cypselae

obovoid-ellipsoid, slightly flattened, faces minutely hairy or papillate (papilliform hairs or papillae ± clavate, not myxogenic);

pappi readily falling (singly or in groups), of 12–20, distinct or basally coherent, barbellate bristles in 1 series.

Inner

(bisexual) florets 1–7;

corollas purple or distally purplish.

Pappus

bristles distinct or basally coherent (falling in groups).

x

= 14.

2n

= 28.

Euchiton

Euchiton sphaericus

Phenology Flowering Jul–Oct.
Habitat Grassy open places in wooded areas, disturbed soils, recent clearings
Elevation 30–700 m (100–2300 ft)
Distribution
from USDA
Australia; New Zealand; New Guinea; e Asia; some species widely naturalized [Introduced in North America]
[BONAP county map]
from FNA
CA; OR; se Asia; Pacific Islands (Hawaii, New Guinea, New Zealand); Australia [Introduced in North America]
[WildflowerSearch map]
[BONAP county map]
Discussion

Species 17 (3 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

In California and Hawaii, plants of Euchiton sphaericus have been identified as Gnaphalium japonicum; annual duration, slender taproots, non-clasping leaf bases, and single bisexual florets establish the correct identity of E. sphaericus. It “varies enormously in length, position and degree of branching, and the branches may be either vegetative or flower-bearing” (D. G. Drury 1972).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Annuals; taprooted; leaf bases not clasping; bracts subtending heads 4–8; heads in globose clusters; bisexual florets 1
E. sphaericus
1. Perennials or biennials; fibrous-rooted; leaf bases subclasping; bracts subtending heads 2–5; heads in hemispheric clusters; bisexual florets 3–7
→ 2
2. Stolons usually present; basal leaves in rosettes at flowering; cauline leaves 2–4(–6), blades linear to oblanceolate, 1–2 cm × 1–2 mm; bracts subtending heads 2–3, not surpassing heads; pistillate florets 40–60
E. gymnocephalus
2. Stolons usually absent; basal leaves withering before flowering; cauline leaves 6–10, blades mostly linear, 3–8 cm × 2–3 mm; bracts subtending heads 3–5, surpassing heads; pistillate florets 80–150
E. involucratus
Source FNA vol. 19, p. 440. Author: Guy L. Nesom. FNA vol. 19, p. 441.
Parent taxa Asteraceae > tribe Gnaphalieae Asteraceae > tribe Gnaphalieae > Euchiton
Sibling taxa
E. gymnocephalus, E. involucratus
Subordinate taxa
E. gymnocephalus, E. involucratus, E. sphaericus
Synonyms Gnaphalium sphaericum
Name authority Cassini: in F. Cuvier, Dict. Sci. Nat. ed. 2, 56: 214. (1828) (Willdenow) Anderberg: Opera Bot. 104: 167. (1991)
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