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brickellbush aster, hairy rayless aster, rayless aster

aster

Habit Perennials 40–90 cm (caudices woody). Perennials, 10–160 cm (usually cespitose, induments usually of stipitate-glandular and smooth-surfaced, curved or twisted woolly hairs, plants with caudices or short rhizomes, roots fibrous).
Stems

erect, woolly or cottony.

ascending or erect, simple, glabrate, puberulent, pilose, cottony, or woolly, eglandular or glandular.

Leaves

mid and distal blades lanceolate to elliptic, 2.5–6 cm × 7–20 mm, abaxial faces glabrous or glabrate, adaxial ± densely woolly to cottony.

cauline; alternate;

sessile (proximal withering by flowering; proximalmost reduced, scalelike);

blades (1-nerved) ovate, elliptic, oblong, lanceolate, or linear (± uniform in size), margins entire, faces glabrate, scabrous, cottony, or woolly, eglandular or stipitate-glandular.

Peduncles

pubescent.

Involucres

turbinate, 7–10 mm.

turbinate-cylindric, turbinate, turbinate-obconic, or campanulate, 10–25 mm diam.

Receptacles

± flat, pitted, epaleate.

Ray florets

0–21 (usually 5, 8, or 13), pistillate, fertile;

corollas violet-purple, purple, pink, or white.

Disc florets

10–35, bisexual, fertile;

corollas yellow, ± ampliate, tubes shorter than funnelform throats, lobes 5, erect or reflexed, triangular;

style-branch appendages lanceolate.

Phyllaries

in 4–6 series (often reddish at margins and apices), linear-oblong to ovate (strongly unequal), apices acute, abaxial faces tomentose to stipitate-glandular.

20–50 in 3–6 series, ± unequal (± appressed, often reddish or purplish at margins and tips), 1-nerved (keeled), ovate, lance-oblong, lanceolate, linear-oblong, or linear, chartaceous at bases, margins sometimes hyaline, especially proximally;

apices acute to obtuse, green, usually puberulent, tomentose, and/or stipitate-glandular, sometimes glabrous.

Heads

3–40 in racemiform to paniculiform arrays.

radiate or discoid, usually in open, racemiform, paniculiform, or corymbiform arrays, sometimes borne singly.

Cypselae

glabrous or pilose;

pappus bristles in 2 series, ± barbellate.

± obconic, flattened, laterally 1–2-ribbed, sometimes with 1–2 additional nerves on each face, glabrous, pilose, or strigose, eglandular;

pappi persistent, of 30–50 whitish to tawny, barbellate or smooth, apically clavate or more conspicuously barbellate bristles in 2(–3) series (outer usually 1 mm or less, sometimes 0, inner 5–10 mm).

Rays

(0–)1–3(–6), violet-purple.

x

= 9.

Eucephalus tomentellus

Eucephalus

Phenology Flowering Jul–Aug.
Habitat Open oak or coniferous woods, forest openings and rocky cliffs
Elevation 1300–2400 m (4300–7900 ft)
Distribution
from FNA
CA; OR
[WildflowerSearch map]
[BONAP county map]
from USDA
North America
[BONAP county map]
Discussion

Eucephalus tomentellus grows in the Siskiyou Mountains of southwestern Oregon and northern California. It may intergrade with E. breweri and E. glabratus.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Species 10 (10 in the flora).

Eucephalus, a relatively well-marked western North American group, has been treated as a section of Aster or as a distinct genus. Recent molecular evidence places Eucephalus, together with the eastern North American Doellingeria, at the base of the North American clade of Astereae.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Ray florets usually 1–4, often 0
→ 2
1. Ray florets commonly 5, 8, or 13+
→ 5
2. Ray florets 0; leaves 5–9 cm, ± glabrous abaxially, glandular adaxially; plants 60–120 cm; open woods, Lane County, Oregon
E. vialis
2. Ray florets usually 1–4; leaves 2–6 cm, hairy; plants 10–100 cm
→ 3
3. Leaves glabrous or nearly so abaxially, moderately to densely hairy adaxially
E. tomentellus
3. Leaves glabrous, eglandular or sparsely glandular on both faces
→ 4
4. Phyllaries subequal
E. breweri
4. Phyllaries strongly unequal
E. glabratus
5. Stems, leaves, and phyllaries glabrous, glaucous; plants 40–160 cm; leaves linear tonarrowly lance-elliptic, 4–10 cm; rays purple
E. glaucescens
5. Stems, leaves, and phyllaries pubescent or glabrate, glandular or not, not glaucous; plants 10–120(–150) cm; leaves elliptic, oblong, lance-ovate, lance-elliptic, lanceolate, linear-oblong or -lanceolate, 1.5–10 cm; rays white, pink, violet, or purple.
→ 6
6. Leaves 5–10 cm, elliptic to lanceolate, glabrous and eglandular, or abaxially ± glandular and/or villous; plants 50–150 cm; rays white to pink
E. engelmannii
6. Leaves 1.5–7 cm, elliptic, elliptic-oblong, oblong, lance-ovate, lance-elliptic, linear-oblong or -lanceolate, glandular or not, scabrous or cottony; plants 10–80 cm; rays white or violet to purple
→ 7
7. Rays white; stems pilose or sparsely to moderately glandular-pubescent
→ 8
7. Rays violet to purple; stems scabrous (to scabrellous) or cottony and/or glandular-pubescent (especially peduncles)
→ 9
8. Phyllaries lance-ovate; Cascade Mountains, Oregon
E. gormanii
8. Phyllaries lance-linear; Olympic Mountains, Washington
E. paucicapitatus
9. Leaves moderately scabrellous (and sometimes glandular) on both faces
E. elegans
9. Leaves sparsely scabrous abaxially, strongly cottony adaxially
E. ledophyllus
Source FNA vol. 20, p. 42. FNA vol. 20, p. 39. Author: Geraldine A. Allen.
Parent taxa Asteraceae > tribe Astereae > Eucephalus Asteraceae > tribe Astereae
Sibling taxa
E. breweri, E. elegans, E. engelmannii, E. glabratus, E. glaucescens, E. gormanii, E. ledophyllus, E. paucicapitatus, E. vialis
Subordinate taxa
E. breweri, E. elegans, E. engelmannii, E. glabratus, E. glaucescens, E. gormanii, E. ledophyllus, E. paucicapitatus, E. tomentellus, E. vialis
Synonyms Sericocarpus tomentellus, Aster brickellioides, Aster tomentellus, E. bicolor, E. brickellioides Aster section E., Aster subsection E.
Name authority (Greene) Greene: Pittonia 3: 55. (1896) Nuttall: Trans. Amer. Philos. Soc., n. s. 7: 298. (1840)
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