Erythranthe washingtonensis |
Erythranthe utahensis |
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John Day or Washington monkeyflower, Washington monkey-flower |
Utah monkeyflower |
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Habit | Annuals, fibrous-rooted or filiform-taprooted. | Perennials, rhizomatous, rooting at nodes, rhizomes filiform. |
Stems | erect to ascending, straight or geniculate at nodes, usually many-branched, terete, 5–25 cm, moderately puberulent-glandular to villous-glandular, hairs 0.1–0.8 mm, flattened, sometimes vitreous, distinctly multicellular, gland-tipped. |
erect, sometimes decumbent-ascending proximally, simple or few-branched, 20–50 cm, glabrous or sparsely stipitate-glandular in inflorescence. |
Leaves | cauline, basal not persistent; petiole 2–14 mm; blade palmately veined, deltate or ovate to ovate-lanceolate, 4–16(–23) × 2–11(–16) mm, base rounded to cuneate or truncate, margins denticulate or entire, apex acute, surfaces moderately puberulent-glandular to villous-glandular, hairs 0.1–0.8 mm, flattened, sometimes vitreous, distinctly multicellular, gland-tipped. |
basal and cauline, even-sized or largest near mid stem; petiole 0 mm or proximalmost 2–10 mm; blade palmately 3–5-veined, orbicular or suborbicular to broadly elliptic, broadly ovate, or depressed-ovate, 20–40(–75) × 12–35(–40) mm, base usually truncate to broadly cuneate, margins entire or subentire to mucronulate, shallowly dentate, or denticulate, apex rounded, surfaces glabrous or glabrate to sparsely stipitate-glandular and sparsely pilose, hairs thin-walled, abaxial often glaucous. |
Flowers | herkogamous, 1–6, from proximal to distal nodes. |
herkogamous, 6–16, from proximal to distal nodes, in a loose raceme, distal bracts becoming much reduced. |
Styles | hispid-hirtellous. |
hirtellous. |
Corollas | yellow with small reddish brown dots, abaxial limb with 2 white patches (abaxial ridges), bilaterally symmetric, strongly bilabiate; tube-throat funnelform, 8–10 mm, exserted beyond calyx margin; limb expanded 7–10 mm, lobes obovate-oblong, apex rounded to rounded-cuneate. |
yellow, abaxial limb prominently darker yellow, sparsely purple-spotted, bilaterally symmetric, weakly bilabiate; tube-throat narrowly funnelform to broadly cylindric, 10–15 mm, exserted 5–8 mm beyond calyx margin; limb expanded 12–20 mm. |
Fruiting pedicels | divergent at nearly right angles, 20–50 mm, densely, minutely stipitate-glandular. |
(25–)40–75 mm, stipitate-glandular to short villous, hairs gland-tipped. |
Fruiting calyces | greenish, ridge-angled, tubular, weakly inflated, 6–8 mm, margins distinctly toothed or lobed, densely, minutely stipitate-glandular, lobes pronounced, erect. |
broadly ovate-cylindric, inflated, sagittally compressed, (10–)11–17(–20) mm, stipitate-glandular or minutely hirtellous or a mixture, hairs sometimes also longer, thin-walled, eglandular or glandular, throat not closing, adaxial lobe slightly longer, triangular-blunt. |
Capsules | included, 5–8.5 mm. |
included, 5–7 mm. |
Anthers | included, glabrous. |
included, glabrous. |
2n | = 32. |
= 28, 30. |
Erythranthe washingtonensis |
Erythranthe utahensis |
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Phenology | Flowering May–Sep. | Flowering (Feb–)May–Aug(–Oct). |
Habitat | Shallow basalt gravel in narrow channels and intermittent streams, sandy stream banks, open slopes, rocky shelves near seeps. | Drainage ditches, springs, seeps, wet meadows, margins of ponds and small streams, marshy areas. |
Elevation | 700–1300 m. (2300–4300 ft.) | 1400–2500 m. (4600–8200 ft.) |
Distribution |
OR
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CA; CO; NV; UT |
Discussion | Erythranthe washingtonensis is considered to be extirpated in Washington by the Washington Natural Heritage Program. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Erythranthe utahensis is characterized by its erect stems, prolifically produced filiform rhizomes, basal leaves short-petiolate to subsessile and cauline sessile, blades suborbicular to broadly ovate or depressed ovate with thin-walled villous-glandular hairs on both surfaces, margins subentire, proximal pedicels elongating to 75 mm, and calyces open at maturity. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 396. | FNA vol. 17, p. 410. |
Parent taxa | Phrymaceae > Erythranthe | Phrymaceae > Erythranthe |
Sibling taxa | ||
Synonyms | Mimulus washingtonensis | Mimulus glabratus subsp. utahensis |
Name authority | (Gandoger) G. L. Nesom: Phytoneuron 2012-39: 39. (2012) — (as washingtoniensis) | (Pennell) G. L. Nesom: Phytoneuron 2012-39: 44. (2012) |
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