FNA: Erythranthe washingtonensis vs. Erythranthe trinitiensis

	Erythranthe washingtonensis	Erythranthe trinitiensis
	John Day or Washington monkeyflower, Washington monkey-flower	pink-margined monkeyflower, Trinity Mountains or pink-margined monkeyflower
Habit	Annuals, fibrous-rooted or filiform-taprooted.	Annuals, shallowly fibrous-rooted.
Stems	erect to ascending, straight or geniculate at nodes, usually many-branched, terete, 5–25 cm, moderately puberulent-glandular to villous-glandular, hairs 0.1–0.8 mm, flattened, sometimes vitreous, distinctly multicellular, gland-tipped.	erect, straight at nodes, simple or branched at base, 5–15 cm, puberulent, hairs gland-tipped, glands often dark.
Leaves	cauline, basal not persistent; petiole 2–14 mm; blade palmately veined, deltate or ovate to ovate-lanceolate, 4–16(–23) × 2–11(–16) mm, base rounded to cuneate or truncate, margins denticulate or entire, apex acute, surfaces moderately puberulent-glandular to villous-glandular, hairs 0.1–0.8 mm, flattened, sometimes vitreous, distinctly multicellular, gland-tipped.	mostly cauline, largest at mid stem; petiole 4–8 mm, 1-veined, not winged; blade palmately 3-veined, ovate to elliptic-ovate, 6–17 × 4–9 mm, base attenuate, margins entire or dentate-serrate, teeth 1–2 per side, shallow, apex acute, surfaces puberulent, hairs gland-tipped, glands often dark, adaxial sometimes sparsely villous-glandular.
Flowers	herkogamous, 1–6, from proximal to distal nodes.	herkogamous, 1–12, from proximal to distal nodes.
Styles	hispid-hirtellous.	glabrous.
Corollas	yellow with small reddish brown dots, abaxial limb with 2 white patches (abaxial ridges), bilaterally symmetric, strongly bilabiate; tube-throat funnelform, 8–10 mm, exserted beyond calyx margin; limb expanded 7–10 mm, lobes obovate-oblong, apex rounded to rounded-cuneate.	yellow and light pink to white, tube-throat yellow (inner and outer surfaces), lobes (limb) pink or white with pink distal borders, palate ridges yellow, throat floor and ridges weakly red-spotted, bilaterally symmetric, weakly bilabiate; tube-throat funnelform, 7–10 mm, exserted beyond calyx margin.
Fruiting pedicels	divergent at nearly right angles, 20–50 mm, densely, minutely stipitate-glandular.	divergent-arcuate, 9–17 mm.
Fruiting calyces	greenish, ridge-angled, tubular, weakly inflated, 6–8 mm, margins distinctly toothed or lobed, densely, minutely stipitate-glandular, lobes pronounced, erect.	oblong-ovoid, distinctly inflated, 8–10 mm, margins distinctly toothed or lobed, puberulent, hairs gland-tipped, glands often dark, lobes pronounced, erect.
Capsules	included, 5–8.5 mm.	included, 6–8 mm.
Anthers	included, glabrous.	included, glabrous.
2n	= 32.

	Erythranthe washingtonensis	Erythranthe trinitiensis
Phenology	Flowering May–Sep.	Flowering Jun–Jul(–Aug).
Habitat	Shallow basalt gravel in narrow channels and intermittent streams, sandy stream banks, open slopes, rocky shelves near seeps.	Seeps over serpentine, wet meadows, roadsides.
Elevation	700–1300 m. (2300–4300 ft.)	1300–2000 m. (4300–6600 ft.)
Distribution	OR [WildflowerSearch map]	CA
Discussion	Erythranthe washingtonensis is considered to be extirpated in Washington by the Washington Natural Heritage Program. (Discussion copyrighted by Flora of North America; reprinted with permission.)	Erythranthe trinitiensis is similar to E. pulsiferae in its narrow leaves, glandular-puberulent vestiture, and weakly bilabiate corollas; it differs in its early-shed basal leaves, cauline leaves with one-veined petioles, and yellow corolla tubes and throats with pink lobes or lobe margins. The species is known only in Humboldt, Siskiyou, and Trinity counties. (Discussion copyrighted by Flora of North America; reprinted with permission.)
Source	FNA vol. 17, p. 396.	FNA vol. 17, p. 399.
Parent taxa	Phrymaceae > Erythranthe	Phrymaceae > Erythranthe
Sibling taxa	E. acutidens, E. alsinoides, E. ampliata, E. androsacea, E. arenaria, E. arenicola, E. arvensis, E. barbata, E. bicolor, E. brachystylis, E. breviflora, E. breweri, E. caespitosa, E. calcicola, E. calciphila, E. cardinalis, E. carsonensis, E. charlestonensis, E. chinatiensis, E. cinnabarina, E. corallina, E. cordata, E. decora, E. dentata, E. diffusa, E. discolor, E. eastwoodiae, E. erubescens, E. exigua, E. filicaulis, E. filicifolia, E. floribunda, E. gemmipara, E. geniculata, E. geyeri, E. glaucescens, E. gracilipes, E. grandis, E. grayi, E. guttata, E. hallii, E. hardhamiae, E. hymenophylla, E. inamoena, E. inconspicua, E. inflatula, E. jungermannioides, E. laciniata, E. latidens, E. lewisii, E. linearifolia, E. marmorata, E. michiganensis, E. microphylla, E. minor, E. montioides, E. moschata, E. nasuta, E. norrisii, E. nudata, E. palmeri, E. pardalis, E. parishii, E. parvula, E. patula, E. percaulis, E. primuloides, E. ptilota, E. pulsiferae, E. purpurea, E. regni, E. rhodopetra, E. rubella, E. scouleri, E. shevockii, E. sierrae, E. suksdorfii, E. taylorii, E. thermalis, E. tilingii, E. trinitiensis, E. unimaculata, E. utahensis, E. verbenacea, E. willisii	E. acutidens, E. alsinoides, E. ampliata, E. androsacea, E. arenaria, E. arenicola, E. arvensis, E. barbata, E. bicolor, E. brachystylis, E. breviflora, E. breweri, E. caespitosa, E. calcicola, E. calciphila, E. cardinalis, E. carsonensis, E. charlestonensis, E. chinatiensis, E. cinnabarina, E. corallina, E. cordata, E. decora, E. dentata, E. diffusa, E. discolor, E. eastwoodiae, E. erubescens, E. exigua, E. filicaulis, E. filicifolia, E. floribunda, E. gemmipara, E. geniculata, E. geyeri, E. glaucescens, E. gracilipes, E. grandis, E. grayi, E. guttata, E. hallii, E. hardhamiae, E. hymenophylla, E. inamoena, E. inconspicua, E. inflatula, E. jungermannioides, E. laciniata, E. latidens, E. lewisii, E. linearifolia, E. marmorata, E. michiganensis, E. microphylla, E. minor, E. montioides, E. moschata, E. nasuta, E. norrisii, E. nudata, E. palmeri, E. pardalis, E. parishii, E. parvula, E. patula, E. percaulis, E. primuloides, E. ptilota, E. pulsiferae, E. purpurea, E. regni, E. rhodopetra, E. rubella, E. scouleri, E. shevockii, E. sierrae, E. suksdorfii, E. taylorii, E. thermalis, E. tilingii, E. unimaculata, E. utahensis, E. verbenacea, E. washingtonensis, E. willisii
Synonyms	Mimulus washingtonensis
Name authority	(Gandoger) G. L. Nesom: Phytoneuron 2012-39: 39. (2012) — (as washingtoniensis)	G. L. Nesom: Phytoneuron 2013-43: 1, figs. 1–3. (2013)
Web links	Local floras: OR, WA Local Web sites: Flora NW, PNW Herbaria WildflowerSearch iNaturalist (observations) LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search	Local floras: CA Local Web sites: CalFlora, CalPhotos WildflowerSearch iNaturalist (observations) LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search

Erythranthe trinitiensis

John Day or Washington monkeyflower, Washington monkey-flower

pink-margined monkeyflower, Trinity Mountains or pink-margined monkeyflower

Habit

Annuals, fibrous-rooted or filiform-taprooted.

Annuals, shallowly fibrous-rooted.

Stems

erect to ascending, straight or geniculate at nodes, usually many-branched, terete, 5–25 cm, moderately puberulent-glandular to villous-glandular, hairs 0.1–0.8 mm, flattened, sometimes vitreous, distinctly multicellular, gland-tipped.

erect, straight at nodes, simple or branched at base, 5–15 cm, puberulent, hairs gland-tipped, glands often dark.

Leaves

cauline, basal not persistent;

petiole 2–14 mm;

blade palmately veined, deltate or ovate to ovate-lanceolate, 4–16(–23) × 2–11(–16) mm, base rounded to cuneate or truncate, margins denticulate or entire, apex acute, surfaces moderately puberulent-glandular to villous-glandular, hairs 0.1–0.8 mm, flattened, sometimes vitreous, distinctly multicellular, gland-tipped.

mostly cauline, largest at mid stem;

petiole 4–8 mm, 1-veined, not winged;

blade palmately 3-veined, ovate to elliptic-ovate, 6–17 × 4–9 mm, base attenuate, margins entire or dentate-serrate, teeth 1–2 per side, shallow, apex acute, surfaces puberulent, hairs gland-tipped, glands often dark, adaxial sometimes sparsely villous-glandular.

Flowers

herkogamous, 1–6, from proximal to distal nodes.

herkogamous, 1–12, from proximal to distal nodes.

Styles

hispid-hirtellous.

glabrous.

Corollas

yellow with small reddish brown dots, abaxial limb with 2 white patches (abaxial ridges), bilaterally symmetric, strongly bilabiate;

tube-throat funnelform, 8–10 mm, exserted beyond calyx margin;

limb expanded 7–10 mm, lobes obovate-oblong, apex rounded to rounded-cuneate.

yellow and light pink to white, tube-throat yellow (inner and outer surfaces), lobes (limb) pink or white with pink distal borders, palate ridges yellow, throat floor and ridges weakly red-spotted, bilaterally symmetric, weakly bilabiate;

tube-throat funnelform, 7–10 mm, exserted beyond calyx margin.

Fruiting pedicels

divergent at nearly right angles, 20–50 mm, densely, minutely stipitate-glandular.

divergent-arcuate, 9–17 mm.

Fruiting calyces

greenish, ridge-angled, tubular, weakly inflated, 6–8 mm, margins distinctly toothed or lobed, densely, minutely stipitate-glandular, lobes pronounced, erect.

oblong-ovoid, distinctly inflated, 8–10 mm, margins distinctly toothed or lobed, puberulent, hairs gland-tipped, glands often dark, lobes pronounced, erect.

Capsules

included, 5–8.5 mm.

included, 6–8 mm.

Anthers

included, glabrous.

= 32.

Erythranthe washingtonensis

Erythranthe trinitiensis

Phenology

Flowering May–Sep.

Flowering Jun–Jul(–Aug).

Habitat

Shallow basalt gravel in narrow channels and intermittent streams, sandy stream banks, open slopes, rocky shelves near seeps.

Seeps over serpentine, wet meadows, roadsides.

Elevation

700–1300 m. (2300–4300 ft.)

1300–2000 m. (4300–6600 ft.)

Distribution

[WildflowerSearch map]

Discussion

Erythranthe washingtonensis is considered to be extirpated in Washington by the Washington Natural Heritage Program.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Erythranthe trinitiensis is similar to E. pulsiferae in its narrow leaves, glandular-puberulent vestiture, and weakly bilabiate corollas; it differs in its early-shed basal leaves, cauline leaves with one-veined petioles, and yellow corolla tubes and throats with pink lobes or lobe margins. The species is known only in Humboldt, Siskiyou, and Trinity counties.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source

FNA vol. 17, p. 396.

FNA vol. 17, p. 399.

Parent taxa

Phrymaceae > Erythranthe

Sibling taxa

E. acutidens, E. alsinoides, E. ampliata, E. androsacea, E. arenaria, E. arenicola, E. arvensis, E. barbata, E. bicolor, E. brachystylis, E. breviflora, E. breweri, E. caespitosa, E. calcicola, E. calciphila, E. cardinalis, E. carsonensis, E. charlestonensis, E. chinatiensis, E. cinnabarina, E. corallina, E. cordata, E. decora, E. dentata, E. diffusa, E. discolor, E. eastwoodiae, E. erubescens, E. exigua, E. filicaulis, E. filicifolia, E. floribunda, E. gemmipara, E. geniculata, E. geyeri, E. glaucescens, E. gracilipes, E. grandis, E. grayi, E. guttata, E. hallii, E. hardhamiae, E. hymenophylla, E. inamoena, E. inconspicua, E. inflatula, E. jungermannioides, E. laciniata, E. latidens, E. lewisii, E. linearifolia, E. marmorata, E. michiganensis, E. microphylla, E. minor, E. montioides, E. moschata, E. nasuta, E. norrisii, E. nudata, E. palmeri, E. pardalis, E. parishii, E. parvula, E. patula, E. percaulis, E. primuloides, E. ptilota, E. pulsiferae, E. purpurea, E. regni, E. rhodopetra, E. rubella, E. scouleri, E. shevockii, E. sierrae, E. suksdorfii, E. taylorii, E. thermalis, E. tilingii, E. trinitiensis, E. unimaculata, E. utahensis, E. verbenacea, E. willisii

E. acutidens, E. alsinoides, E. ampliata, E. androsacea, E. arenaria, E. arenicola, E. arvensis, E. barbata, E. bicolor, E. brachystylis, E. breviflora, E. breweri, E. caespitosa, E. calcicola, E. calciphila, E. cardinalis, E. carsonensis, E. charlestonensis, E. chinatiensis, E. cinnabarina, E. corallina, E. cordata, E. decora, E. dentata, E. diffusa, E. discolor, E. eastwoodiae, E. erubescens, E. exigua, E. filicaulis, E. filicifolia, E. floribunda, E. gemmipara, E. geniculata, E. geyeri, E. glaucescens, E. gracilipes, E. grandis, E. grayi, E. guttata, E. hallii, E. hardhamiae, E. hymenophylla, E. inamoena, E. inconspicua, E. inflatula, E. jungermannioides, E. laciniata, E. latidens, E. lewisii, E. linearifolia, E. marmorata, E. michiganensis, E. microphylla, E. minor, E. montioides, E. moschata, E. nasuta, E. norrisii, E. nudata, E. palmeri, E. pardalis, E. parishii, E. parvula, E. patula, E. percaulis, E. primuloides, E. ptilota, E. pulsiferae, E. purpurea, E. regni, E. rhodopetra, E. rubella, E. scouleri, E. shevockii, E. sierrae, E. suksdorfii, E. taylorii, E. thermalis, E. tilingii, E. unimaculata, E. utahensis, E. verbenacea, E. washingtonensis, E. willisii

Synonyms

Mimulus washingtonensis

Name authority

(Gandoger) G. L. Nesom: Phytoneuron 2012-39: 39. (2012) — (as washingtoniensis)

G. L. Nesom: Phytoneuron 2013-43: 1, figs. 1–3. (2013)

Web links

Unless otherwise noted, all data comes from the online version of the Flora of North America, who have made most of their content available via a Creative Commons license (CC BY 4.0).

Choose a data source: Flora of North America, OregonFlora, UW Burke Herbarium

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Erythranthe washingtonensis

Erythranthe trinitiensis

Erythranthe washingtonensis

Erythranthe trinitiensis