Erythranthe washingtonensis |
Erythranthe taylorii |
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John Day or Washington monkeyflower, Washington monkey-flower |
Shasta limestone monkeyflower, Taylor's or Shasta limestone monkeyflower |
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Habit | Annuals, fibrous-rooted or filiform-taprooted. | Annuals, filiform-taprooted. |
Stems | erect to ascending, straight or geniculate at nodes, usually many-branched, terete, 5–25 cm, moderately puberulent-glandular to villous-glandular, hairs 0.1–0.8 mm, flattened, sometimes vitreous, distinctly multicellular, gland-tipped. |
erect, straight at nodes, simple or few-branched from base, 5–10 cm, sparsely eglandular-villous proximally, becoming sparsely short stipitate-glandular distally. |
Leaves | cauline, basal not persistent; petiole 2–14 mm; blade palmately veined, deltate or ovate to ovate-lanceolate, 4–16(–23) × 2–11(–16) mm, base rounded to cuneate or truncate, margins denticulate or entire, apex acute, surfaces moderately puberulent-glandular to villous-glandular, hairs 0.1–0.8 mm, flattened, sometimes vitreous, distinctly multicellular, gland-tipped. |
usually cauline, basal not persistent, largest at mid stem or basal and mid stem to nearly even-sized; petiole 3–5(–8) mm; blade often purple adaxially, palmately 3–5-veined, broadly ovate to elliptic-ovate or ovate-lanceolate, 4–20 × 4–12 mm, base rounded to truncate, margins serrate-dentate, teeth 2–4 per side, shallow, apex rounded to obtuse or acute, surfaces: distals moderately short-stipitate-glandular. |
Flowers | herkogamous, 1–6, from proximal to distal nodes. |
herkogamous, sometimes plesiogamous, 2–6(–8), from proximal to distal nodes. |
Styles | hispid-hirtellous. |
glabrous. |
Corollas | yellow with small reddish brown dots, abaxial limb with 2 white patches (abaxial ridges), bilaterally symmetric, strongly bilabiate; tube-throat funnelform, 8–10 mm, exserted beyond calyx margin; limb expanded 7–10 mm, lobes obovate-oblong, apex rounded to rounded-cuneate. |
yellow, throat ceiling sometimes red-spotted or -lined, abaxial limb yellow or with 1 or 2 red splotches, bilaterally symmetric, bilabiate; tube-throat funnelform, 5–7 mm, exserted beyond calyx margin. |
Fruiting pedicels | divergent at nearly right angles, 20–50 mm, densely, minutely stipitate-glandular. |
divergent to arcuate-divergent, 6–13 mm. |
Fruiting calyces | greenish, ridge-angled, tubular, weakly inflated, 6–8 mm, margins distinctly toothed or lobed, densely, minutely stipitate-glandular, lobes pronounced, erect. |
wing-angled, tubular-campanulate, 4–5 mm, margins distinctly toothed or lobed, densely invested with tiny, waxy-white, eglandular, papillose hairs between angles, lobes pronounced, erect. |
Capsules | included, 5–8.5 mm. |
included, 3–4 mm. |
Anthers | included, glabrous. |
included, glabrous. |
2n | = 32. |
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Erythranthe washingtonensis |
Erythranthe taylorii |
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Phenology | Flowering May–Sep. | Flowering Feb–May. |
Habitat | Shallow basalt gravel in narrow channels and intermittent streams, sandy stream banks, open slopes, rocky shelves near seeps. | Crevices in limestone cliff faces and outcrops. |
Elevation | 700–1300 m. (2300–4300 ft.) | 900–1100 m. (3000–3600 ft.) |
Distribution |
OR
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CA
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Discussion | Erythranthe washingtonensis is considered to be extirpated in Washington by the Washington Natural Heritage Program. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Erythranthe taylorii is known only from the Shasta Lake region of northwestern Shasta County. Its broad, distinctly bilabiate corollas and ovate leaf blades with palmate venation are similar to those of species of the northern group of sect. Mimulosma, the "Columbia River clade" (J. B. Whittall et al. 2006) of Idaho, Montana, Oregon, Washington, and Wyoming, particularly to the Idaho endemic E. ampliata. Erythranthe taylorii is distinct from E. ampliata in its larger, papillose calyces, shorter fruiting pedicels, corollas with shorter tube-throats, and shorter capsules. Considerable corolla color variation exists in E. taylorii in the occurrence and density of red dots and lines on the throat ceiling and larger red splotches on the abaxial limb. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 396. | FNA vol. 17, p. 398. |
Parent taxa | Phrymaceae > Erythranthe | Phrymaceae > Erythranthe |
Sibling taxa | ||
Synonyms | Mimulus washingtonensis | |
Name authority | (Gandoger) G. L. Nesom: Phytoneuron 2012-39: 39. (2012) — (as washingtoniensis) | G. L. Nesom: Phytoneuron 2013-43: 6, figs. 5–7. (2013) — (as taylori) |
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