FNA: Erythranthe washingtonensis vs. Erythranthe parvula

	Erythranthe washingtonensis	Erythranthe parvula
	John Day or Washington monkeyflower, Washington monkey-flower	Southwestern mat monkeyflower
Habit	Annuals, fibrous-rooted or filiform-taprooted.	Perennials, rhizomatous, sometimes rooting at nodes, mat-forming.
Stems	erect to ascending, straight or geniculate at nodes, usually many-branched, terete, 5–25 cm, moderately puberulent-glandular to villous-glandular, hairs 0.1–0.8 mm, flattened, sometimes vitreous, distinctly multicellular, gland-tipped.	procumbent, branched, 5–15 cm, stipitate-glandular distally.
Leaves	cauline, basal not persistent; petiole 2–14 mm; blade palmately veined, deltate or ovate to ovate-lanceolate, 4–16(–23) × 2–11(–16) mm, base rounded to cuneate or truncate, margins denticulate or entire, apex acute, surfaces moderately puberulent-glandular to villous-glandular, hairs 0.1–0.8 mm, flattened, sometimes vitreous, distinctly multicellular, gland-tipped.	cauline; petiole 1–4 mm; blade palmately 3-veined, ovate to orbicular-ovate or depressed-ovate, 3–11 × 3–9 mm, base truncate to cuneate, margins shallowly denticulate to dentate, teeth 3–5 per side, apex acute to obtuse, surfaces villous-hirsute, hairs whitish, thickened, flattened, stiff, gland-tipped.
Flowers	herkogamous, 1–6, from proximal to distal nodes.	plesiogamous, 2–8, axillary at distal nodes.
Styles	hispid-hirtellous.	glabrous.
Corollas	yellow with small reddish brown dots, abaxial limb with 2 white patches (abaxial ridges), bilaterally symmetric, strongly bilabiate; tube-throat funnelform, 8–10 mm, exserted beyond calyx margin; limb expanded 7–10 mm, lobes obovate-oblong, apex rounded to rounded-cuneate.	yellow, red-dotted, bilaterally symmetric, bilabiate; tube-throat funnelform, 6–8 mm, exserted beyond calyx margin; limb expanded 4–6 mm, abaxial limb spreading, lobes fimbriate.
Fruiting pedicels	divergent at nearly right angles, 20–50 mm, densely, minutely stipitate-glandular.	7–15 mm, minutely stipitate-glandular.
Fruiting calyces	greenish, ridge-angled, tubular, weakly inflated, 6–8 mm, margins distinctly toothed or lobed, densely, minutely stipitate-glandular, lobes pronounced, erect.	nodding 45–90º, 5-lobed, cylindric-ovoid, inflated, sagittally compressed, 4–5 mm, villous-hirsute, throat closing.
Capsules	included, 5–8.5 mm.	included, 3–4 mm.
Anthers	included, glabrous.	included, glabrous.
2n	= 32.	= 32.

	Erythranthe washingtonensis	Erythranthe parvula
Phenology	Flowering May–Sep.	Flowering Apr–Sep.
Habitat	Shallow basalt gravel in narrow channels and intermittent streams, sandy stream banks, open slopes, rocky shelves near seeps.	Wet vertical rock faces, ledges, and rocky slopes, seepy wash banks.
Elevation	700–1300 m. (2300–4300 ft.)	500–2400(–3400) m. (1600–7900(–11200) ft.)
Distribution	OR [WildflowerSearch map]	AZ; NM; Mexico (Sonora)
Discussion	Erythranthe washingtonensis is considered to be extirpated in Washington by the Washington Natural Heritage Program. (Discussion copyrighted by Flora of North America; reprinted with permission.)	All monkeyflowers with laciniate-lobed corollas have generally been identified as Mimulus dentilobus B. L. Robinson & Fernald, but these occur as three, morphologically distinct, widely allopatric population systems, each of which is regarded as a separate species (G. L. Nesom 2012g). Erythranthe dentiloba (B. L. Robinson & Fernald) G. L. Nesom, which is endemic to Mexico, is the only one of the three with an allogamous breeding system. Erythranthe parvula is restricted to Arizona and New Mexico except for one locality in northern Sonora, Mexico. Erythranthe chinatiensis is the third species of the group. (Discussion copyrighted by Flora of North America; reprinted with permission.)
Source	FNA vol. 17, p. 396.	FNA vol. 17, p. 424.
Parent taxa	Phrymaceae > Erythranthe	Phrymaceae > Erythranthe
Sibling taxa	E. acutidens, E. alsinoides, E. ampliata, E. androsacea, E. arenaria, E. arenicola, E. arvensis, E. barbata, E. bicolor, E. brachystylis, E. breviflora, E. breweri, E. caespitosa, E. calcicola, E. calciphila, E. cardinalis, E. carsonensis, E. charlestonensis, E. chinatiensis, E. cinnabarina, E. corallina, E. cordata, E. decora, E. dentata, E. diffusa, E. discolor, E. eastwoodiae, E. erubescens, E. exigua, E. filicaulis, E. filicifolia, E. floribunda, E. gemmipara, E. geniculata, E. geyeri, E. glaucescens, E. gracilipes, E. grandis, E. grayi, E. guttata, E. hallii, E. hardhamiae, E. hymenophylla, E. inamoena, E. inconspicua, E. inflatula, E. jungermannioides, E. laciniata, E. latidens, E. lewisii, E. linearifolia, E. marmorata, E. michiganensis, E. microphylla, E. minor, E. montioides, E. moschata, E. nasuta, E. norrisii, E. nudata, E. palmeri, E. pardalis, E. parishii, E. parvula, E. patula, E. percaulis, E. primuloides, E. ptilota, E. pulsiferae, E. purpurea, E. regni, E. rhodopetra, E. rubella, E. scouleri, E. shevockii, E. sierrae, E. suksdorfii, E. taylorii, E. thermalis, E. tilingii, E. trinitiensis, E. unimaculata, E. utahensis, E. verbenacea, E. willisii	E. acutidens, E. alsinoides, E. ampliata, E. androsacea, E. arenaria, E. arenicola, E. arvensis, E. barbata, E. bicolor, E. brachystylis, E. breviflora, E. breweri, E. caespitosa, E. calcicola, E. calciphila, E. cardinalis, E. carsonensis, E. charlestonensis, E. chinatiensis, E. cinnabarina, E. corallina, E. cordata, E. decora, E. dentata, E. diffusa, E. discolor, E. eastwoodiae, E. erubescens, E. exigua, E. filicaulis, E. filicifolia, E. floribunda, E. gemmipara, E. geniculata, E. geyeri, E. glaucescens, E. gracilipes, E. grandis, E. grayi, E. guttata, E. hallii, E. hardhamiae, E. hymenophylla, E. inamoena, E. inconspicua, E. inflatula, E. jungermannioides, E. laciniata, E. latidens, E. lewisii, E. linearifolia, E. marmorata, E. michiganensis, E. microphylla, E. minor, E. montioides, E. moschata, E. nasuta, E. norrisii, E. nudata, E. palmeri, E. pardalis, E. parishii, E. patula, E. percaulis, E. primuloides, E. ptilota, E. pulsiferae, E. purpurea, E. regni, E. rhodopetra, E. rubella, E. scouleri, E. shevockii, E. sierrae, E. suksdorfii, E. taylorii, E. thermalis, E. tilingii, E. trinitiensis, E. unimaculata, E. utahensis, E. verbenacea, E. washingtonensis, E. willisii
Synonyms	Mimulus washingtonensis	Mimulus parvulus
Name authority	(Gandoger) G. L. Nesom: Phytoneuron 2012-39: 39. (2012) — (as washingtoniensis)	(Wooton & Standley) G. L. Nesom: Phytoneuron 2012-39: 44. (2012)
Web links	Local floras: OR, WA Local Web sites: Flora NW, PNW Herbaria WildflowerSearch iNaturalist (observations) LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search	WildflowerSearch iNaturalist (observations) LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search

Erythranthe parvula

John Day or Washington monkeyflower, Washington monkey-flower

Southwestern mat monkeyflower

Habit

Annuals, fibrous-rooted or filiform-taprooted.

Perennials, rhizomatous, sometimes rooting at nodes, mat-forming.

Stems

erect to ascending, straight or geniculate at nodes, usually many-branched, terete, 5–25 cm, moderately puberulent-glandular to villous-glandular, hairs 0.1–0.8 mm, flattened, sometimes vitreous, distinctly multicellular, gland-tipped.

procumbent, branched, 5–15 cm, stipitate-glandular distally.

Leaves

cauline, basal not persistent;

petiole 2–14 mm;

blade palmately veined, deltate or ovate to ovate-lanceolate, 4–16(–23) × 2–11(–16) mm, base rounded to cuneate or truncate, margins denticulate or entire, apex acute, surfaces moderately puberulent-glandular to villous-glandular, hairs 0.1–0.8 mm, flattened, sometimes vitreous, distinctly multicellular, gland-tipped.

cauline;

petiole 1–4 mm;

blade palmately 3-veined, ovate to orbicular-ovate or depressed-ovate, 3–11 × 3–9 mm, base truncate to cuneate, margins shallowly denticulate to dentate, teeth 3–5 per side, apex acute to obtuse, surfaces villous-hirsute, hairs whitish, thickened, flattened, stiff, gland-tipped.

Flowers

herkogamous, 1–6, from proximal to distal nodes.

plesiogamous, 2–8, axillary at distal nodes.

Styles

hispid-hirtellous.

glabrous.

Corollas

yellow with small reddish brown dots, abaxial limb with 2 white patches (abaxial ridges), bilaterally symmetric, strongly bilabiate;

tube-throat funnelform, 8–10 mm, exserted beyond calyx margin;

limb expanded 7–10 mm, lobes obovate-oblong, apex rounded to rounded-cuneate.

yellow, red-dotted, bilaterally symmetric, bilabiate;

tube-throat funnelform, 6–8 mm, exserted beyond calyx margin;

limb expanded 4–6 mm, abaxial limb spreading, lobes fimbriate.

Fruiting pedicels

divergent at nearly right angles, 20–50 mm, densely, minutely stipitate-glandular.

7–15 mm, minutely stipitate-glandular.

Fruiting calyces

greenish, ridge-angled, tubular, weakly inflated, 6–8 mm, margins distinctly toothed or lobed, densely, minutely stipitate-glandular, lobes pronounced, erect.

nodding 45–90º, 5-lobed, cylindric-ovoid, inflated, sagittally compressed, 4–5 mm, villous-hirsute, throat closing.

Capsules

included, 5–8.5 mm.

included, 3–4 mm.

Anthers

included, glabrous.

= 32.

Erythranthe washingtonensis

Erythranthe parvula

Phenology

Flowering May–Sep.

Flowering Apr–Sep.

Habitat

Shallow basalt gravel in narrow channels and intermittent streams, sandy stream banks, open slopes, rocky shelves near seeps.

Wet vertical rock faces, ledges, and rocky slopes, seepy wash banks.

Elevation

700–1300 m. (2300–4300 ft.)

500–2400(–3400) m. (1600–7900(–11200) ft.)

Distribution

[WildflowerSearch map]

AZ; NM; Mexico (Sonora)

Discussion

Erythranthe washingtonensis is considered to be extirpated in Washington by the Washington Natural Heritage Program.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

All monkeyflowers with laciniate-lobed corollas have generally been identified as Mimulus dentilobus B. L. Robinson & Fernald, but these occur as three, morphologically distinct, widely allopatric population systems, each of which is regarded as a separate species (G. L. Nesom 2012g). Erythranthe dentiloba (B. L. Robinson & Fernald) G. L. Nesom, which is endemic to Mexico, is the only one of the three with an allogamous breeding system. Erythranthe parvula is restricted to Arizona and New Mexico except for one locality in northern Sonora, Mexico. Erythranthe chinatiensis is the third species of the group.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source

FNA vol. 17, p. 396.

FNA vol. 17, p. 424.

Parent taxa

Phrymaceae > Erythranthe

Sibling taxa

E. acutidens, E. alsinoides, E. ampliata, E. androsacea, E. arenaria, E. arenicola, E. arvensis, E. barbata, E. bicolor, E. brachystylis, E. breviflora, E. breweri, E. caespitosa, E. calcicola, E. calciphila, E. cardinalis, E. carsonensis, E. charlestonensis, E. chinatiensis, E. cinnabarina, E. corallina, E. cordata, E. decora, E. dentata, E. diffusa, E. discolor, E. eastwoodiae, E. erubescens, E. exigua, E. filicaulis, E. filicifolia, E. floribunda, E. gemmipara, E. geniculata, E. geyeri, E. glaucescens, E. gracilipes, E. grandis, E. grayi, E. guttata, E. hallii, E. hardhamiae, E. hymenophylla, E. inamoena, E. inconspicua, E. inflatula, E. jungermannioides, E. laciniata, E. latidens, E. lewisii, E. linearifolia, E. marmorata, E. michiganensis, E. microphylla, E. minor, E. montioides, E. moschata, E. nasuta, E. norrisii, E. nudata, E. palmeri, E. pardalis, E. parishii, E. parvula, E. patula, E. percaulis, E. primuloides, E. ptilota, E. pulsiferae, E. purpurea, E. regni, E. rhodopetra, E. rubella, E. scouleri, E. shevockii, E. sierrae, E. suksdorfii, E. taylorii, E. thermalis, E. tilingii, E. trinitiensis, E. unimaculata, E. utahensis, E. verbenacea, E. willisii

E. acutidens, E. alsinoides, E. ampliata, E. androsacea, E. arenaria, E. arenicola, E. arvensis, E. barbata, E. bicolor, E. brachystylis, E. breviflora, E. breweri, E. caespitosa, E. calcicola, E. calciphila, E. cardinalis, E. carsonensis, E. charlestonensis, E. chinatiensis, E. cinnabarina, E. corallina, E. cordata, E. decora, E. dentata, E. diffusa, E. discolor, E. eastwoodiae, E. erubescens, E. exigua, E. filicaulis, E. filicifolia, E. floribunda, E. gemmipara, E. geniculata, E. geyeri, E. glaucescens, E. gracilipes, E. grandis, E. grayi, E. guttata, E. hallii, E. hardhamiae, E. hymenophylla, E. inamoena, E. inconspicua, E. inflatula, E. jungermannioides, E. laciniata, E. latidens, E. lewisii, E. linearifolia, E. marmorata, E. michiganensis, E. microphylla, E. minor, E. montioides, E. moschata, E. nasuta, E. norrisii, E. nudata, E. palmeri, E. pardalis, E. parishii, E. patula, E. percaulis, E. primuloides, E. ptilota, E. pulsiferae, E. purpurea, E. regni, E. rhodopetra, E. rubella, E. scouleri, E. shevockii, E. sierrae, E. suksdorfii, E. taylorii, E. thermalis, E. tilingii, E. trinitiensis, E. unimaculata, E. utahensis, E. verbenacea, E. washingtonensis, E. willisii

Synonyms

Mimulus washingtonensis

Mimulus parvulus

Name authority

(Gandoger) G. L. Nesom: Phytoneuron 2012-39: 39. (2012) — (as washingtoniensis)

(Wooton & Standley) G. L. Nesom: Phytoneuron 2012-39: 44. (2012)

Web links

Unless otherwise noted, all data comes from the online version of the Flora of North America, who have made most of their content available via a Creative Commons license (CC BY 4.0).

Choose a data source: Flora of North America, OregonFlora, UW Burke Herbarium

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Flora of North America species comparison

Erythranthe washingtonensis

Erythranthe parvula

Erythranthe washingtonensis

Erythranthe parvula