Erythranthe washingtonensis |
Erythranthe laciniata |
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John Day or Washington monkeyflower, Washington monkey-flower |
cut-leaf monkeyflower |
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Habit | Annuals, fibrous-rooted or filiform-taprooted. | Annuals, slender-taprooted or fibrous-rooted. |
Stems | erect to ascending, straight or geniculate at nodes, usually many-branched, terete, 5–25 cm, moderately puberulent-glandular to villous-glandular, hairs 0.1–0.8 mm, flattened, sometimes vitreous, distinctly multicellular, gland-tipped. |
erect, simple or branched from base, 3–38 cm, glabrous or sparsely hirtellous, finely villosulous-glandular above nodes. |
Leaves | cauline, basal not persistent; petiole 2–14 mm; blade palmately veined, deltate or ovate to ovate-lanceolate, 4–16(–23) × 2–11(–16) mm, base rounded to cuneate or truncate, margins denticulate or entire, apex acute, surfaces moderately puberulent-glandular to villous-glandular, hairs 0.1–0.8 mm, flattened, sometimes vitreous, distinctly multicellular, gland-tipped. |
cauline, basal deciduous by flowering; petiole 1–35 mm, distals 0 mm; blade 1-veined or palmately 3-veined, elliptic to elliptic-obovate, oblanceolate, or oblong, 3–55 mm, longer than wide, base attenuate, margins narrowly pinnately lobed or dissected, sometimes merely shallowly toothed, apex acute to obtuse, surfaces glabrate. |
Flowers | herkogamous, 1–6, from proximal to distal nodes. |
plesiogamous, 2–8, from medial to distal nodes, chasmogamous, sometimes cleistogamous. |
Styles | hispid-hirtellous. |
glabrous. |
Corollas | yellow with small reddish brown dots, abaxial limb with 2 white patches (abaxial ridges), bilaterally symmetric, strongly bilabiate; tube-throat funnelform, 8–10 mm, exserted beyond calyx margin; limb expanded 7–10 mm, lobes obovate-oblong, apex rounded to rounded-cuneate. |
yellow, throat red-spotted, abaxial limb of larger usually with 1 large red splotch, bilaterally symmetric, ± bilabiate; tube-throat funnelform, 4–6 mm, exserted 1–2 mm beyond calyx margin; limb expanded 5–6 mm. |
Fruiting pedicels | divergent at nearly right angles, 20–50 mm, densely, minutely stipitate-glandular. |
nodding 30–140º at calyx base, 5–25 mm. |
Fruiting calyces | greenish, ridge-angled, tubular, weakly inflated, 6–8 mm, margins distinctly toothed or lobed, densely, minutely stipitate-glandular, lobes pronounced, erect. |
red-spotted, cylindric-campanulate, inflated, sagittally compressed, 8–10 mm, glabrate, throat closing, lobes ca. equal size or adaxial slightly longer. |
Capsules | included, 5–8.5 mm. |
included, stipitate, 5–7 mm. |
Anthers | included, glabrous. |
included, glabrous. |
2n | = 32. |
= 28. |
Erythranthe washingtonensis |
Erythranthe laciniata |
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Phenology | Flowering May–Sep. | Flowering Apr–Jul(–Aug). |
Habitat | Shallow basalt gravel in narrow channels and intermittent streams, sandy stream banks, open slopes, rocky shelves near seeps. | Cracks, depressions, and seeps in granite outcrops, ledges, talus and scree, rocky streamsides, rocky slopes, roadsides, intermittent drainages. |
Elevation | 700–1300 m. (2300–4300 ft.) | 900–2300(–3300) m. (3000–7500(–10800) ft.) |
Distribution |
OR
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CA
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Discussion | Erythranthe washingtonensis is considered to be extirpated in Washington by the Washington Natural Heritage Program. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Erythranthe laciniata is known from Amador County south to Kern County. As in Erythranthe nasuta, the adaxial calyx lobe in E. laciniata tends to be narrowly lanceolate to triangular (noselike) and perceptibly falcate, curving slightly upward both in flower and in fruit. The adaxial lobe is not so prominently protruding as it often is in E. nasuta. Corolla size is variable in Erythranthe laciniata, but the size of those with an open throat (versus much reduced in size and apparently cleistogamous) is not strongly correlated with size of the individual plant, and all on one plant are about the same size (compare with E. nasuta). Corollas on some plants, however, are all or nearly all greatly reduced and apparently cleistogamous. Fertilization in even the larger corollas apparently is autogamous; the anther pairs are slightly separated or equal in level, and the stigma is in the middle of the anthers or at the level of the adaxial pair. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 396. | FNA vol. 17, p. 419. |
Parent taxa | Phrymaceae > Erythranthe | Phrymaceae > Erythranthe |
Sibling taxa | ||
Synonyms | Mimulus washingtonensis | Mimulus laciniatus, M. eisenii |
Name authority | (Gandoger) G. L. Nesom: Phytoneuron 2012-39: 39. (2012) — (as washingtoniensis) | (A. Gray) G. L. Nesom: Phytoneuron 2012-39: 44. (2012) |
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