Erythranthe washingtonensis |
Erythranthe inamoena |
|
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John Day or Washington monkeyflower, Washington monkey-flower |
Texas monkeyflower |
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Habit | Annuals, fibrous-rooted or filiform-taprooted. | Annuals, fibrous-rooted, rooting at proximal nodes, sometimes forming matlike colonies. |
Stems | erect to ascending, straight or geniculate at nodes, usually many-branched, terete, 5–25 cm, moderately puberulent-glandular to villous-glandular, hairs 0.1–0.8 mm, flattened, sometimes vitreous, distinctly multicellular, gland-tipped. |
decumbent to decumbent-ascending basally, becoming fully erect at least in inflorescence, simple, sometimes few-branched from proximal nodes, usually distinctly fistulose, 10–30 cm, glabrous. |
Leaves | cauline, basal not persistent; petiole 2–14 mm; blade palmately veined, deltate or ovate to ovate-lanceolate, 4–16(–23) × 2–11(–16) mm, base rounded to cuneate or truncate, margins denticulate or entire, apex acute, surfaces moderately puberulent-glandular to villous-glandular, hairs 0.1–0.8 mm, flattened, sometimes vitreous, distinctly multicellular, gland-tipped. |
basal and cauline, basal sometimes deciduous by flowering; petiole: basal and proximals to mid cauline 10–70 mm, distals 0 mm; blade palmately veined, ovate to broadly ovate or elliptic-ovate, becoming subreniform distally, (5–)15–35(–60) mm, distal closely paired, auriculate-subclasping, base truncate to subcordate, margins dentate-serrate to shallowly dentate, teeth 5–11 per side, apex obtuse to rounded, surfaces glabrous. |
Flowers | herkogamous, 1–6, from proximal to distal nodes. |
plesiogamous, (6–)8–18(–24), often produced from all nodes, loosely to densely racemose, chasmogamous. |
Styles | hispid-hirtellous. |
glabrous. |
Corollas | yellow with small reddish brown dots, abaxial limb with 2 white patches (abaxial ridges), bilaterally symmetric, strongly bilabiate; tube-throat funnelform, 8–10 mm, exserted beyond calyx margin; limb expanded 7–10 mm, lobes obovate-oblong, apex rounded to rounded-cuneate. |
yellow, red-spotted, bilaterally symmetric, bilabiate; tube-throat cylindric, 7–11 mm, exserted (1–)2–3(–4) mm beyond calyx margin; limb slightly expanded. |
Fruiting pedicels | divergent at nearly right angles, 20–50 mm, densely, minutely stipitate-glandular. |
9–20 mm, glabrous, rarely sparsely stipitate-glandular. |
Fruiting calyces | greenish, ridge-angled, tubular, weakly inflated, 6–8 mm, margins distinctly toothed or lobed, densely, minutely stipitate-glandular, lobes pronounced, erect. |
purple-spotted, sometimes greenish, broadly cylindric-campanulate, inflated, sagittally compressed, 7–11 mm, glabrous, rarely minutely scabrous-hirtellous or sparsely stipitate-glandular, throat not closing, abaxial lobe slightly upcurving 10–45º, spreading 45º, or sometimes deflexed 40º. |
Capsules | included, 5–8.5 mm. |
included, 4.5–6 mm. |
Anthers | included, glabrous. |
included, glabrous. |
2n | = 32. |
= 60. |
Erythranthe washingtonensis |
Erythranthe inamoena |
|
Phenology | Flowering May–Sep. | Flowering Jan–Apr(–Sep). |
Habitat | Shallow basalt gravel in narrow channels and intermittent streams, sandy stream banks, open slopes, rocky shelves near seeps. | Edge of seeps and creeks, mud or gravel, shallow running water, wet crevices, canyon drainages. |
Elevation | 700–1300 m. (2300–4300 ft.) | 100–2400 m. (300–7900 ft.) |
Distribution |
OR
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TX; Mexico (Chihuahua, Coahuila) |
Discussion | Erythranthe washingtonensis is considered to be extirpated in Washington by the Washington Natural Heritage Program. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Erythranthe inamoena is distinctive in its completely glabrous herbage, small corollas, flowers in racemes mostly at distal nodes and with reduced bracts, short and open-throated fruiting calyces, erect and fistulose stems, and apparent annual duration (fibrous-rooted but usually rooting at proximal cauline nodes). Presumably because of its autogamous reproduction, Erythranthe inamoena has been confused with E. cordata, especially in the trans-Pecos region of Texas, where the two are sympatric. In contrast to E. cordata, E. inamoena usually has glabrous pedicels and calyces, flowers often produced from all nodes, smaller calyces [(7–)8–11 mm] that do not close at maturity, and corollas with a shorter tube-throat (7–11 mm). Some populations in Brewster, Presidio, and Val Verde counties are identified here as Erythranthe inamoena (based on proximal-to-distal distribution of flowers and the short mature calyces with open throat) but have sparsely stipitate-glandular pedicels and calyces. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 396. | FNA vol. 17, p. 407. |
Parent taxa | Phrymaceae > Erythranthe | Phrymaceae > Erythranthe |
Sibling taxa | ||
Synonyms | Mimulus washingtonensis | Mimulus inamoenus, M. jamesii var. texensis |
Name authority | (Gandoger) G. L. Nesom: Phytoneuron 2012-39: 39. (2012) — (as washingtoniensis) | (Greene) G. L. Nesom: Phytoneuron 2012-39: 44. (2012) |
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