Erythranthe washingtonensis |
Erythranthe hymenophylla |
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John Day or Washington monkeyflower, Washington monkey-flower |
thin-sepal monkeyflower |
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Habit | Annuals, fibrous-rooted or filiform-taprooted. | Annuals, filiform-taprooted. |
Stems | erect to ascending, straight or geniculate at nodes, usually many-branched, terete, 5–25 cm, moderately puberulent-glandular to villous-glandular, hairs 0.1–0.8 mm, flattened, sometimes vitreous, distinctly multicellular, gland-tipped. |
prostrate to ascending-erect, sharply bent at basal nodes, simple or few-branched, 5–25 cm, glandular-puberulent to glandular-villous, hairs 0.1–0.8 mm, vitreous, flattened, multicellular, gland-tipped. |
Leaves | cauline, basal not persistent; petiole 2–14 mm; blade palmately veined, deltate or ovate to ovate-lanceolate, 4–16(–23) × 2–11(–16) mm, base rounded to cuneate or truncate, margins denticulate or entire, apex acute, surfaces moderately puberulent-glandular to villous-glandular, hairs 0.1–0.8 mm, flattened, sometimes vitreous, distinctly multicellular, gland-tipped. |
basal and cauline, largest at mid stem; petiole 6–30 mm; blade pinnately veined, broadly lanceolate to ovate, 10–35 × 10–30 mm, distinctly membranous, base cuneate to shallowly cordate, margins coarsely dentate to shallowly denticulate or entire, apex acute to obtuse, surfaces glandular-puberulent to glandular-villous, hairs 0.1–0.8 mm, vitreous, flattened, multicellular, gland-tipped, glandular. |
Flowers | herkogamous, 1–6, from proximal to distal nodes. |
herkogamous, 1–6, from proximal to distal nodes. |
Styles | hispid-hirtellous. |
glabrous. |
Corollas | yellow with small reddish brown dots, abaxial limb with 2 white patches (abaxial ridges), bilaterally symmetric, strongly bilabiate; tube-throat funnelform, 8–10 mm, exserted beyond calyx margin; limb expanded 7–10 mm, lobes obovate-oblong, apex rounded to rounded-cuneate. |
light yellow, throat and abaxial lobes red- or purple-spotted, sometimes with small white patches, bilaterally symmetric, weakly bilabiate; tube-throat funnelform, 10–14 mm, exserted beyond calyx margin; lobes obovate-oblong, apex rounded to truncate or notched. |
Fruiting pedicels | divergent at nearly right angles, 20–50 mm, densely, minutely stipitate-glandular. |
divergent at right angles from stem, usually closely paired, 10–45 mm, negatively phototropic, causing capsules to be pressed against a cliff face or crevice at time of dehiscence, glandular-puberulent to glandular-villous, hairs 0.1–0.8 mm, vitreous, flattened, multicellular, gland-tipped. |
Fruiting calyces | greenish, ridge-angled, tubular, weakly inflated, 6–8 mm, margins distinctly toothed or lobed, densely, minutely stipitate-glandular, lobes pronounced, erect. |
angled, tubular-campanulate, slightly inflated, 5–7 mm, margins distinctly toothed or lobed, sparsely stipitate-glandular, lobes pronounced, erect. |
Capsules | included, 5–8.5 mm. |
included, 3–6 mm. |
Anthers | included, glabrous. |
included, glabrous. |
2n | = 32. |
= 32. |
Erythranthe washingtonensis |
Erythranthe hymenophylla |
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Phenology | Flowering May–Sep. | Flowering Apr–Aug(–Sep). |
Habitat | Shallow basalt gravel in narrow channels and intermittent streams, sandy stream banks, open slopes, rocky shelves near seeps. | Steep, seasonally moist, basalt cliffs with west or southwest exposure, mesic coniferous forests. |
Elevation | 700–1300 m. (2300–4300 ft.) | 800–1300 m. (2600–4300 ft.) |
Distribution |
OR
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ID; MT; OR |
Discussion | Erythranthe washingtonensis is considered to be extirpated in Washington by the Washington Natural Heritage Program. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
In the protologue, R. J. Meinke observed that plants of Erythranthe hymenophylla have reflexed fruiting pedicels that increase seed dispersal back onto the vertical cliff wall, the characteristic habitat of the species. The hanging habit of E. hymenophylla is reflected in a sharp (90º to 180º) bend in the basal nodes and the long pedicels that are closely paired and divergent in parallel at about right angles from the stem. The species also is characterized by it very short calyx to corolla length, relatively short capsules, and large seeds. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 396. | FNA vol. 17, p. 397. |
Parent taxa | Phrymaceae > Erythranthe | Phrymaceae > Erythranthe |
Sibling taxa | ||
Synonyms | Mimulus washingtonensis | Mimulus hymenophyllus |
Name authority | (Gandoger) G. L. Nesom: Phytoneuron 2012-39: 39. (2012) — (as washingtoniensis) | (Meinke) G. L. Nesom: Phytoneuron 2012-39: 38. (2012) |
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