FNA: Erythranthe washingtonensis vs. Erythranthe gracilipes

	Erythranthe washingtonensis	Erythranthe gracilipes
	John Day or Washington monkeyflower, Washington monkey-flower	slender-stalk monkeyflower
Habit	Annuals, fibrous-rooted or filiform-taprooted.	Annuals, taprooted.
Stems	erect to ascending, straight or geniculate at nodes, usually many-branched, terete, 5–25 cm, moderately puberulent-glandular to villous-glandular, hairs 0.1–0.8 mm, flattened, sometimes vitreous, distinctly multicellular, gland-tipped.	erect, simple or branched, 5–15 cm, sparsely glandular-puberulent.
Leaves	cauline, basal not persistent; petiole 2–14 mm; blade palmately veined, deltate or ovate to ovate-lanceolate, 4–16(–23) × 2–11(–16) mm, base rounded to cuneate or truncate, margins denticulate or entire, apex acute, surfaces moderately puberulent-glandular to villous-glandular, hairs 0.1–0.8 mm, flattened, sometimes vitreous, distinctly multicellular, gland-tipped.	cauline, basal not persistent; petiole 0 mm; blade 1-veined or palmately 3-veined (in broader ones), linear to lanceolate, (3–)7–13 × 1–3 mm, base attenuate, margins entire, sometimes toothed, apex acute to obtuse, surfaces sparsely glandular-puberulent.
Flowers	herkogamous, 1–6, from proximal to distal nodes.	herkogamous, 1–15, from distal or medial to distal nodes.
Styles	hispid-hirtellous.	glabrous.
Corollas	yellow with small reddish brown dots, abaxial limb with 2 white patches (abaxial ridges), bilaterally symmetric, strongly bilabiate; tube-throat funnelform, 8–10 mm, exserted beyond calyx margin; limb expanded 7–10 mm, lobes obovate-oblong, apex rounded to rounded-cuneate.	pink to rose lavender, throat deep pink to purple with 2 yellow longitudinal ridges, bilaterally symmetric, strongly bilabiate, 2 adaxial lobes much reduced, smaller than 3 abaxials; tube-throat cylindric, 7–10 mm, exserted beyond calyx margin; limb expanded 5–9 mm, abaxial lobes rounded and shallowly 2-fid, abaxial limb glabrous or sparsely bearded.
Fruiting pedicels	divergent at nearly right angles, 20–50 mm, densely, minutely stipitate-glandular.	8–30 mm.
Fruiting calyces	greenish, ridge-angled, tubular, weakly inflated, 6–8 mm, margins distinctly toothed or lobed, densely, minutely stipitate-glandular, lobes pronounced, erect.	becoming red to straw colored, campanulate, 4–7 mm, margins distinctly toothed or lobed, sparsely glandular-puberulent, ribs thickened, lobes pronounced, erect, margins ciliate.
Capsules	included, 5–8.5 mm.	included, 4–5 mm.
Anthers	included, glabrous.	included, glabrous.
2n	= 32.

	Erythranthe washingtonensis	Erythranthe gracilipes
Phenology	Flowering May–Sep.	Flowering Apr–May.
Habitat	Shallow basalt gravel in narrow channels and intermittent streams, sandy stream banks, open slopes, rocky shelves near seeps.	Open areas in thin decomposed granite soils, often on edges of large granite boulders.
Elevation	700–1300 m. (2300–4300 ft.)	500–1300 m. (1600–4300 ft.)
Distribution	OR [WildflowerSearch map]	CA [WildflowerSearch map]
Discussion	Erythranthe washingtonensis is considered to be extirpated in Washington by the Washington Natural Heritage Program. (Discussion copyrighted by Flora of North America; reprinted with permission.)	Erythranthe gracilipes is endemic to the foothills of the central Sierra Nevada in Fresno, Madera, and Mariposa counties and apparently is most abundant after fire. (Discussion copyrighted by Flora of North America; reprinted with permission.)
Source	FNA vol. 17, p. 396.	FNA vol. 17, p. 385.
Parent taxa	Phrymaceae > Erythranthe	Phrymaceae > Erythranthe
Sibling taxa	E. acutidens, E. alsinoides, E. ampliata, E. androsacea, E. arenaria, E. arenicola, E. arvensis, E. barbata, E. bicolor, E. brachystylis, E. breviflora, E. breweri, E. caespitosa, E. calcicola, E. calciphila, E. cardinalis, E. carsonensis, E. charlestonensis, E. chinatiensis, E. cinnabarina, E. corallina, E. cordata, E. decora, E. dentata, E. diffusa, E. discolor, E. eastwoodiae, E. erubescens, E. exigua, E. filicaulis, E. filicifolia, E. floribunda, E. gemmipara, E. geniculata, E. geyeri, E. glaucescens, E. gracilipes, E. grandis, E. grayi, E. guttata, E. hallii, E. hardhamiae, E. hymenophylla, E. inamoena, E. inconspicua, E. inflatula, E. jungermannioides, E. laciniata, E. latidens, E. lewisii, E. linearifolia, E. marmorata, E. michiganensis, E. microphylla, E. minor, E. montioides, E. moschata, E. nasuta, E. norrisii, E. nudata, E. palmeri, E. pardalis, E. parishii, E. parvula, E. patula, E. percaulis, E. primuloides, E. ptilota, E. pulsiferae, E. purpurea, E. regni, E. rhodopetra, E. rubella, E. scouleri, E. shevockii, E. sierrae, E. suksdorfii, E. taylorii, E. thermalis, E. tilingii, E. trinitiensis, E. unimaculata, E. utahensis, E. verbenacea, E. willisii	E. acutidens, E. alsinoides, E. ampliata, E. androsacea, E. arenaria, E. arenicola, E. arvensis, E. barbata, E. bicolor, E. brachystylis, E. breviflora, E. breweri, E. caespitosa, E. calcicola, E. calciphila, E. cardinalis, E. carsonensis, E. charlestonensis, E. chinatiensis, E. cinnabarina, E. corallina, E. cordata, E. decora, E. dentata, E. diffusa, E. discolor, E. eastwoodiae, E. erubescens, E. exigua, E. filicaulis, E. filicifolia, E. floribunda, E. gemmipara, E. geniculata, E. geyeri, E. glaucescens, E. grandis, E. grayi, E. guttata, E. hallii, E. hardhamiae, E. hymenophylla, E. inamoena, E. inconspicua, E. inflatula, E. jungermannioides, E. laciniata, E. latidens, E. lewisii, E. linearifolia, E. marmorata, E. michiganensis, E. microphylla, E. minor, E. montioides, E. moschata, E. nasuta, E. norrisii, E. nudata, E. palmeri, E. pardalis, E. parishii, E. parvula, E. patula, E. percaulis, E. primuloides, E. ptilota, E. pulsiferae, E. purpurea, E. regni, E. rhodopetra, E. rubella, E. scouleri, E. shevockii, E. sierrae, E. suksdorfii, E. taylorii, E. thermalis, E. tilingii, E. trinitiensis, E. unimaculata, E. utahensis, E. verbenacea, E. washingtonensis, E. willisii
Synonyms	Mimulus washingtonensis	Mimulus gracilipes
Name authority	(Gandoger) G. L. Nesom: Phytoneuron 2012-39: 39. (2012) — (as washingtoniensis)	(B. L. Robinson) N. S. Fraga: Phytoneuron 2012-39: 35. (2012)
Web links	Local floras: OR, WA Local Web sites: Flora NW, PNW Herbaria WildflowerSearch iNaturalist (observations) LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search	Local floras: CA Local Web sites: CalFlora, CalPhotos WildflowerSearch iNaturalist (observations) LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search

Erythranthe gracilipes

John Day or Washington monkeyflower, Washington monkey-flower

slender-stalk monkeyflower

Habit

Annuals, fibrous-rooted or filiform-taprooted.

Annuals, taprooted.

Stems

erect to ascending, straight or geniculate at nodes, usually many-branched, terete, 5–25 cm, moderately puberulent-glandular to villous-glandular, hairs 0.1–0.8 mm, flattened, sometimes vitreous, distinctly multicellular, gland-tipped.

erect, simple or branched, 5–15 cm, sparsely glandular-puberulent.

Leaves

cauline, basal not persistent;

petiole 2–14 mm;

blade palmately veined, deltate or ovate to ovate-lanceolate, 4–16(–23) × 2–11(–16) mm, base rounded to cuneate or truncate, margins denticulate or entire, apex acute, surfaces moderately puberulent-glandular to villous-glandular, hairs 0.1–0.8 mm, flattened, sometimes vitreous, distinctly multicellular, gland-tipped.

cauline, basal not persistent;

petiole 0 mm;

blade 1-veined or palmately 3-veined (in broader ones), linear to lanceolate, (3–)7–13 × 1–3 mm, base attenuate, margins entire, sometimes toothed, apex acute to obtuse, surfaces sparsely glandular-puberulent.

Flowers

herkogamous, 1–6, from proximal to distal nodes.

herkogamous, 1–15, from distal or medial to distal nodes.

Styles

hispid-hirtellous.

glabrous.

Corollas

yellow with small reddish brown dots, abaxial limb with 2 white patches (abaxial ridges), bilaterally symmetric, strongly bilabiate;

tube-throat funnelform, 8–10 mm, exserted beyond calyx margin;

limb expanded 7–10 mm, lobes obovate-oblong, apex rounded to rounded-cuneate.

pink to rose lavender, throat deep pink to purple with 2 yellow longitudinal ridges, bilaterally symmetric, strongly bilabiate, 2 adaxial lobes much reduced, smaller than 3 abaxials;

tube-throat cylindric, 7–10 mm, exserted beyond calyx margin;

limb expanded 5–9 mm, abaxial lobes rounded and shallowly 2-fid, abaxial limb glabrous or sparsely bearded.

Fruiting pedicels

divergent at nearly right angles, 20–50 mm, densely, minutely stipitate-glandular.

8–30 mm.

Fruiting calyces

greenish, ridge-angled, tubular, weakly inflated, 6–8 mm, margins distinctly toothed or lobed, densely, minutely stipitate-glandular, lobes pronounced, erect.

becoming red to straw colored, campanulate, 4–7 mm, margins distinctly toothed or lobed, sparsely glandular-puberulent, ribs thickened, lobes pronounced, erect, margins ciliate.

Capsules

included, 5–8.5 mm.

included, 4–5 mm.

Anthers

included, glabrous.

= 32.

Erythranthe washingtonensis

Erythranthe gracilipes

Phenology

Flowering May–Sep.

Flowering Apr–May.

Habitat

Shallow basalt gravel in narrow channels and intermittent streams, sandy stream banks, open slopes, rocky shelves near seeps.

Open areas in thin decomposed granite soils, often on edges of large granite boulders.

Elevation

700–1300 m. (2300–4300 ft.)

500–1300 m. (1600–4300 ft.)

Distribution

[WildflowerSearch map]

Discussion

Erythranthe washingtonensis is considered to be extirpated in Washington by the Washington Natural Heritage Program.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Erythranthe gracilipes is endemic to the foothills of the central Sierra Nevada in Fresno, Madera, and Mariposa counties and apparently is most abundant after fire.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source

FNA vol. 17, p. 396.

FNA vol. 17, p. 385.

Parent taxa

Phrymaceae > Erythranthe

Sibling taxa

E. acutidens, E. alsinoides, E. ampliata, E. androsacea, E. arenaria, E. arenicola, E. arvensis, E. barbata, E. bicolor, E. brachystylis, E. breviflora, E. breweri, E. caespitosa, E. calcicola, E. calciphila, E. cardinalis, E. carsonensis, E. charlestonensis, E. chinatiensis, E. cinnabarina, E. corallina, E. cordata, E. decora, E. dentata, E. diffusa, E. discolor, E. eastwoodiae, E. erubescens, E. exigua, E. filicaulis, E. filicifolia, E. floribunda, E. gemmipara, E. geniculata, E. geyeri, E. glaucescens, E. gracilipes, E. grandis, E. grayi, E. guttata, E. hallii, E. hardhamiae, E. hymenophylla, E. inamoena, E. inconspicua, E. inflatula, E. jungermannioides, E. laciniata, E. latidens, E. lewisii, E. linearifolia, E. marmorata, E. michiganensis, E. microphylla, E. minor, E. montioides, E. moschata, E. nasuta, E. norrisii, E. nudata, E. palmeri, E. pardalis, E. parishii, E. parvula, E. patula, E. percaulis, E. primuloides, E. ptilota, E. pulsiferae, E. purpurea, E. regni, E. rhodopetra, E. rubella, E. scouleri, E. shevockii, E. sierrae, E. suksdorfii, E. taylorii, E. thermalis, E. tilingii, E. trinitiensis, E. unimaculata, E. utahensis, E. verbenacea, E. willisii

E. acutidens, E. alsinoides, E. ampliata, E. androsacea, E. arenaria, E. arenicola, E. arvensis, E. barbata, E. bicolor, E. brachystylis, E. breviflora, E. breweri, E. caespitosa, E. calcicola, E. calciphila, E. cardinalis, E. carsonensis, E. charlestonensis, E. chinatiensis, E. cinnabarina, E. corallina, E. cordata, E. decora, E. dentata, E. diffusa, E. discolor, E. eastwoodiae, E. erubescens, E. exigua, E. filicaulis, E. filicifolia, E. floribunda, E. gemmipara, E. geniculata, E. geyeri, E. glaucescens, E. grandis, E. grayi, E. guttata, E. hallii, E. hardhamiae, E. hymenophylla, E. inamoena, E. inconspicua, E. inflatula, E. jungermannioides, E. laciniata, E. latidens, E. lewisii, E. linearifolia, E. marmorata, E. michiganensis, E. microphylla, E. minor, E. montioides, E. moschata, E. nasuta, E. norrisii, E. nudata, E. palmeri, E. pardalis, E. parishii, E. parvula, E. patula, E. percaulis, E. primuloides, E. ptilota, E. pulsiferae, E. purpurea, E. regni, E. rhodopetra, E. rubella, E. scouleri, E. shevockii, E. sierrae, E. suksdorfii, E. taylorii, E. thermalis, E. tilingii, E. trinitiensis, E. unimaculata, E. utahensis, E. verbenacea, E. washingtonensis, E. willisii

Synonyms

Mimulus washingtonensis

Mimulus gracilipes

Name authority

(Gandoger) G. L. Nesom: Phytoneuron 2012-39: 39. (2012) — (as washingtoniensis)

(B. L. Robinson) N. S. Fraga: Phytoneuron 2012-39: 35. (2012)

Web links

Unless otherwise noted, all data comes from the online version of the Flora of North America, who have made most of their content available via a Creative Commons license (CC BY 4.0).

Choose a data source: Flora of North America, OregonFlora, UW Burke Herbarium

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Flora of North America species comparison

Erythranthe washingtonensis

Erythranthe gracilipes

Erythranthe washingtonensis

Erythranthe gracilipes