FNA: Erythranthe washingtonensis vs. Erythranthe filicaulis

	Erythranthe washingtonensis	Erythranthe filicaulis
	John Day or Washington monkeyflower, Washington monkey-flower	Hetch Hetchy monkey-flower, slender-stem monkeyflower
Habit	Annuals, fibrous-rooted or filiform-taprooted.	Annuals, taprooted.
Stems	erect to ascending, straight or geniculate at nodes, usually many-branched, terete, 5–25 cm, moderately puberulent-glandular to villous-glandular, hairs 0.1–0.8 mm, flattened, sometimes vitreous, distinctly multicellular, gland-tipped.	erect, simple or branched at basal nodes, 4–30 cm, densely glandular-puberulent, hairs 0.05–0.1 mm, gland-tipped.
Leaves	cauline, basal not persistent; petiole 2–14 mm; blade palmately veined, deltate or ovate to ovate-lanceolate, 4–16(–23) × 2–11(–16) mm, base rounded to cuneate or truncate, margins denticulate or entire, apex acute, surfaces moderately puberulent-glandular to villous-glandular, hairs 0.1–0.8 mm, flattened, sometimes vitreous, distinctly multicellular, gland-tipped.	cauline, basal usually not persistent; petiole 0 mm; blade 1-veined or palmately 3-veined, linear to oblanceolate, 6–15 × (1–)2–3 mm, base attenuate, margins entire to remotely mucronulate, teeth 1–3 per side, apex acute, surfaces densely glandular-puberulent, hairs 0.05–0.1 mm, gland-tipped.
Flowers	herkogamous, 1–6, from proximal to distal nodes.	herkogamous, 2–8, from medial to distal nodes.
Styles	hispid-hirtellous.	glabrous.
Corollas	yellow with small reddish brown dots, abaxial limb with 2 white patches (abaxial ridges), bilaterally symmetric, strongly bilabiate; tube-throat funnelform, 8–10 mm, exserted beyond calyx margin; limb expanded 7–10 mm, lobes obovate-oblong, apex rounded to rounded-cuneate.	pink to purple, throat dark red-purple, palate ridges yellow, bilaterally or nearly radially symmetric, weakly bilabiate or nearly regular; tube-throat cylindric-funnelform, 8–12(–17) mm, exserted beyond calyx margin; limb expanded 10–14 mm, lobes notched, palate villous.
Fruiting pedicels	divergent at nearly right angles, 20–50 mm, densely, minutely stipitate-glandular.	9–30 mm, densely glandular-puberulent, hairs 0.05–0.1 mm, gland-tipped.
Fruiting calyces	greenish, ridge-angled, tubular, weakly inflated, 6–8 mm, margins distinctly toothed or lobed, densely, minutely stipitate-glandular, lobes pronounced, erect.	reddish brown-spotted, strongly angled, cylindric-campanulate, 7–9 mm, margins distinctly toothed or lobed, densely glandular-puberulent, hairs 0.05–0.1 mm, gland-tipped, ribs corky, lobes pronounced, spreading.
Capsules	included, 5–8.5 mm.	included, 5–7 mm.
Anthers	included, glabrous.	included, hirtellous.
2n	= 32.	= 16.

	Erythranthe washingtonensis	Erythranthe filicaulis
Phenology	Flowering May–Sep.	Flowering May–Jul.
Habitat	Shallow basalt gravel in narrow channels and intermittent streams, sandy stream banks, open slopes, rocky shelves near seeps.	Moist open areas on gentle slopes, meadows, roadsides, gravelly soils.
Elevation	700–1300 m. (2300–4300 ft.)	800–1800 m. (2600–5900 ft.)
Distribution	OR [WildflowerSearch map]	CA [WildflowerSearch map]
Discussion	Erythranthe washingtonensis is considered to be extirpated in Washington by the Washington Natural Heritage Program. (Discussion copyrighted by Flora of North America; reprinted with permission.)	Erythranthe filicaulis is known only for certain from Mariposa and Tuolumne counties. Kappler 1691 (UCLA) from Madera County also may be this species. (Discussion copyrighted by Flora of North America; reprinted with permission.)
Source	FNA vol. 17, p. 396.	FNA vol. 17, p. 391.
Parent taxa	Phrymaceae > Erythranthe	Phrymaceae > Erythranthe
Sibling taxa	E. acutidens, E. alsinoides, E. ampliata, E. androsacea, E. arenaria, E. arenicola, E. arvensis, E. barbata, E. bicolor, E. brachystylis, E. breviflora, E. breweri, E. caespitosa, E. calcicola, E. calciphila, E. cardinalis, E. carsonensis, E. charlestonensis, E. chinatiensis, E. cinnabarina, E. corallina, E. cordata, E. decora, E. dentata, E. diffusa, E. discolor, E. eastwoodiae, E. erubescens, E. exigua, E. filicaulis, E. filicifolia, E. floribunda, E. gemmipara, E. geniculata, E. geyeri, E. glaucescens, E. gracilipes, E. grandis, E. grayi, E. guttata, E. hallii, E. hardhamiae, E. hymenophylla, E. inamoena, E. inconspicua, E. inflatula, E. jungermannioides, E. laciniata, E. latidens, E. lewisii, E. linearifolia, E. marmorata, E. michiganensis, E. microphylla, E. minor, E. montioides, E. moschata, E. nasuta, E. norrisii, E. nudata, E. palmeri, E. pardalis, E. parishii, E. parvula, E. patula, E. percaulis, E. primuloides, E. ptilota, E. pulsiferae, E. purpurea, E. regni, E. rhodopetra, E. rubella, E. scouleri, E. shevockii, E. sierrae, E. suksdorfii, E. taylorii, E. thermalis, E. tilingii, E. trinitiensis, E. unimaculata, E. utahensis, E. verbenacea, E. willisii	E. acutidens, E. alsinoides, E. ampliata, E. androsacea, E. arenaria, E. arenicola, E. arvensis, E. barbata, E. bicolor, E. brachystylis, E. breviflora, E. breweri, E. caespitosa, E. calcicola, E. calciphila, E. cardinalis, E. carsonensis, E. charlestonensis, E. chinatiensis, E. cinnabarina, E. corallina, E. cordata, E. decora, E. dentata, E. diffusa, E. discolor, E. eastwoodiae, E. erubescens, E. exigua, E. filicifolia, E. floribunda, E. gemmipara, E. geniculata, E. geyeri, E. glaucescens, E. gracilipes, E. grandis, E. grayi, E. guttata, E. hallii, E. hardhamiae, E. hymenophylla, E. inamoena, E. inconspicua, E. inflatula, E. jungermannioides, E. laciniata, E. latidens, E. lewisii, E. linearifolia, E. marmorata, E. michiganensis, E. microphylla, E. minor, E. montioides, E. moschata, E. nasuta, E. norrisii, E. nudata, E. palmeri, E. pardalis, E. parishii, E. parvula, E. patula, E. percaulis, E. primuloides, E. ptilota, E. pulsiferae, E. purpurea, E. regni, E. rhodopetra, E. rubella, E. scouleri, E. shevockii, E. sierrae, E. suksdorfii, E. taylorii, E. thermalis, E. tilingii, E. trinitiensis, E. unimaculata, E. utahensis, E. verbenacea, E. washingtonensis, E. willisii
Synonyms	Mimulus washingtonensis	Mimulus filicaulis, M. biolettii
Name authority	(Gandoger) G. L. Nesom: Phytoneuron 2012-39: 39. (2012) — (as washingtoniensis)	(S. Watson) G. L. Nesom & N. S. Fraga: Phytoneuron 2012-39: 36. (2012)
Web links	Local floras: OR, WA Local Web sites: Flora NW, PNW Herbaria WildflowerSearch iNaturalist (observations) LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search	Local floras: CA Local Web sites: CalFlora, CalPhotos WildflowerSearch iNaturalist (observations) LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search

Erythranthe filicaulis

John Day or Washington monkeyflower, Washington monkey-flower

Hetch Hetchy monkey-flower, slender-stem monkeyflower

Habit

Annuals, fibrous-rooted or filiform-taprooted.

Annuals, taprooted.

Stems

erect to ascending, straight or geniculate at nodes, usually many-branched, terete, 5–25 cm, moderately puberulent-glandular to villous-glandular, hairs 0.1–0.8 mm, flattened, sometimes vitreous, distinctly multicellular, gland-tipped.

erect, simple or branched at basal nodes, 4–30 cm, densely glandular-puberulent, hairs 0.05–0.1 mm, gland-tipped.

Leaves

cauline, basal not persistent;

petiole 2–14 mm;

blade palmately veined, deltate or ovate to ovate-lanceolate, 4–16(–23) × 2–11(–16) mm, base rounded to cuneate or truncate, margins denticulate or entire, apex acute, surfaces moderately puberulent-glandular to villous-glandular, hairs 0.1–0.8 mm, flattened, sometimes vitreous, distinctly multicellular, gland-tipped.

cauline, basal usually not persistent;

petiole 0 mm;

blade 1-veined or palmately 3-veined, linear to oblanceolate, 6–15 × (1–)2–3 mm, base attenuate, margins entire to remotely mucronulate, teeth 1–3 per side, apex acute, surfaces densely glandular-puberulent, hairs 0.05–0.1 mm, gland-tipped.

Flowers

herkogamous, 1–6, from proximal to distal nodes.

herkogamous, 2–8, from medial to distal nodes.

Styles

hispid-hirtellous.

glabrous.

Corollas

yellow with small reddish brown dots, abaxial limb with 2 white patches (abaxial ridges), bilaterally symmetric, strongly bilabiate;

tube-throat funnelform, 8–10 mm, exserted beyond calyx margin;

limb expanded 7–10 mm, lobes obovate-oblong, apex rounded to rounded-cuneate.

pink to purple, throat dark red-purple, palate ridges yellow, bilaterally or nearly radially symmetric, weakly bilabiate or nearly regular;

tube-throat cylindric-funnelform, 8–12(–17) mm, exserted beyond calyx margin;

limb expanded 10–14 mm, lobes notched, palate villous.

Fruiting pedicels

divergent at nearly right angles, 20–50 mm, densely, minutely stipitate-glandular.

9–30 mm, densely glandular-puberulent, hairs 0.05–0.1 mm, gland-tipped.

Fruiting calyces

greenish, ridge-angled, tubular, weakly inflated, 6–8 mm, margins distinctly toothed or lobed, densely, minutely stipitate-glandular, lobes pronounced, erect.

reddish brown-spotted, strongly angled, cylindric-campanulate, 7–9 mm, margins distinctly toothed or lobed, densely glandular-puberulent, hairs 0.05–0.1 mm, gland-tipped, ribs corky, lobes pronounced, spreading.

Capsules

included, 5–8.5 mm.

included, 5–7 mm.

Anthers

included, glabrous.

included, hirtellous.

= 32.

= 16.

Erythranthe washingtonensis

Erythranthe filicaulis

Phenology

Flowering May–Sep.

Flowering May–Jul.

Habitat

Shallow basalt gravel in narrow channels and intermittent streams, sandy stream banks, open slopes, rocky shelves near seeps.

Moist open areas on gentle slopes, meadows, roadsides, gravelly soils.

Elevation

700–1300 m. (2300–4300 ft.)

800–1800 m. (2600–5900 ft.)

Distribution

[WildflowerSearch map]

Discussion

Erythranthe washingtonensis is considered to be extirpated in Washington by the Washington Natural Heritage Program.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Erythranthe filicaulis is known only for certain from Mariposa and Tuolumne counties. Kappler 1691 (UCLA) from Madera County also may be this species.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source

FNA vol. 17, p. 396.

FNA vol. 17, p. 391.

Parent taxa

Phrymaceae > Erythranthe

Sibling taxa

E. acutidens, E. alsinoides, E. ampliata, E. androsacea, E. arenaria, E. arenicola, E. arvensis, E. barbata, E. bicolor, E. brachystylis, E. breviflora, E. breweri, E. caespitosa, E. calcicola, E. calciphila, E. cardinalis, E. carsonensis, E. charlestonensis, E. chinatiensis, E. cinnabarina, E. corallina, E. cordata, E. decora, E. dentata, E. diffusa, E. discolor, E. eastwoodiae, E. erubescens, E. exigua, E. filicaulis, E. filicifolia, E. floribunda, E. gemmipara, E. geniculata, E. geyeri, E. glaucescens, E. gracilipes, E. grandis, E. grayi, E. guttata, E. hallii, E. hardhamiae, E. hymenophylla, E. inamoena, E. inconspicua, E. inflatula, E. jungermannioides, E. laciniata, E. latidens, E. lewisii, E. linearifolia, E. marmorata, E. michiganensis, E. microphylla, E. minor, E. montioides, E. moschata, E. nasuta, E. norrisii, E. nudata, E. palmeri, E. pardalis, E. parishii, E. parvula, E. patula, E. percaulis, E. primuloides, E. ptilota, E. pulsiferae, E. purpurea, E. regni, E. rhodopetra, E. rubella, E. scouleri, E. shevockii, E. sierrae, E. suksdorfii, E. taylorii, E. thermalis, E. tilingii, E. trinitiensis, E. unimaculata, E. utahensis, E. verbenacea, E. willisii

E. acutidens, E. alsinoides, E. ampliata, E. androsacea, E. arenaria, E. arenicola, E. arvensis, E. barbata, E. bicolor, E. brachystylis, E. breviflora, E. breweri, E. caespitosa, E. calcicola, E. calciphila, E. cardinalis, E. carsonensis, E. charlestonensis, E. chinatiensis, E. cinnabarina, E. corallina, E. cordata, E. decora, E. dentata, E. diffusa, E. discolor, E. eastwoodiae, E. erubescens, E. exigua, E. filicifolia, E. floribunda, E. gemmipara, E. geniculata, E. geyeri, E. glaucescens, E. gracilipes, E. grandis, E. grayi, E. guttata, E. hallii, E. hardhamiae, E. hymenophylla, E. inamoena, E. inconspicua, E. inflatula, E. jungermannioides, E. laciniata, E. latidens, E. lewisii, E. linearifolia, E. marmorata, E. michiganensis, E. microphylla, E. minor, E. montioides, E. moschata, E. nasuta, E. norrisii, E. nudata, E. palmeri, E. pardalis, E. parishii, E. parvula, E. patula, E. percaulis, E. primuloides, E. ptilota, E. pulsiferae, E. purpurea, E. regni, E. rhodopetra, E. rubella, E. scouleri, E. shevockii, E. sierrae, E. suksdorfii, E. taylorii, E. thermalis, E. tilingii, E. trinitiensis, E. unimaculata, E. utahensis, E. verbenacea, E. washingtonensis, E. willisii

Synonyms

Mimulus washingtonensis

Mimulus filicaulis, M. biolettii

Name authority

(Gandoger) G. L. Nesom: Phytoneuron 2012-39: 39. (2012) — (as washingtoniensis)

(S. Watson) G. L. Nesom & N. S. Fraga: Phytoneuron 2012-39: 36. (2012)

Web links

Unless otherwise noted, all data comes from the online version of the Flora of North America, who have made most of their content available via a Creative Commons license (CC BY 4.0).

Choose a data source: Flora of North America, OregonFlora, UW Burke Herbarium

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Flora of North America species comparison

Erythranthe washingtonensis

Erythranthe filicaulis

Erythranthe washingtonensis

Erythranthe filicaulis