FNA: Erythranthe washingtonensis vs. Erythranthe discolor

	Erythranthe washingtonensis	Erythranthe discolor
	John Day or Washington monkeyflower, Washington monkey-flower	party-color monkeyflower, two-color monkeyflower
Habit	Annuals, fibrous-rooted or filiform-taprooted.	Annuals, taprooted.
Stems	erect to ascending, straight or geniculate at nodes, usually many-branched, terete, 5–25 cm, moderately puberulent-glandular to villous-glandular, hairs 0.1–0.8 mm, flattened, sometimes vitreous, distinctly multicellular, gland-tipped.	erect, simple or branched from basal nodes, 5–12 cm, sparsely glandular-pubescent, internodes elongate, distinct.
Leaves	cauline, basal not persistent; petiole 2–14 mm; blade palmately veined, deltate or ovate to ovate-lanceolate, 4–16(–23) × 2–11(–16) mm, base rounded to cuneate or truncate, margins denticulate or entire, apex acute, surfaces moderately puberulent-glandular to villous-glandular, hairs 0.1–0.8 mm, flattened, sometimes vitreous, distinctly multicellular, gland-tipped.	cauline, basal not persistent; petiole 0–2 mm; blade 1-veined or palmately 3-veined (in broader ones), linear to lanceolate or ovate, 5–15 × 1–4 mm, base attenuate, margins entire, apex acute, surfaces sparsely glandular-pubescent.
Flowers	herkogamous, 1–6, from proximal to distal nodes.	herkogamous, 1–8, from distal or medial to distal nodes.
Styles	hispid-hirtellous.	glabrous.
Corollas	yellow with small reddish brown dots, abaxial limb with 2 white patches (abaxial ridges), bilaterally symmetric, strongly bilabiate; tube-throat funnelform, 8–10 mm, exserted beyond calyx margin; limb expanded 7–10 mm, lobes obovate-oblong, apex rounded to rounded-cuneate.	yellow and tube-throat adaxial surface red-tinged, abaxial limb red-spotted or deep pink to purple and abaxial limb with 2 yellow ridges, bilaterally symmetric, strongly bilabiate; tube-throat funnelform, 9–13 mm, exserted beyond calyx margin; limb expanded 7–15 mm, lobes notched, adaxial limb glabrous or sparsely bearded.
Fruiting pedicels	divergent at nearly right angles, 20–50 mm, densely, minutely stipitate-glandular.	erect to ascending, 6–11(–32) mm.
Fruiting calyces	greenish, ridge-angled, tubular, weakly inflated, 6–8 mm, margins distinctly toothed or lobed, densely, minutely stipitate-glandular, lobes pronounced, erect.	sometimes red-dotted on ribs, campanulate, 4–8 mm, margins distinctly toothed or lobed, sparsely glandular-pubescent, ribs weak, lobes pronounced, erect, margins glabrous.
Capsules	included, 5–8.5 mm.	included, 5–8 mm.
Anthers	included, glabrous.	included, glabrous.
2n	= 32.

	Erythranthe washingtonensis	Erythranthe discolor
Phenology	Flowering May–Sep.	Flowering May–Jun.
Habitat	Shallow basalt gravel in narrow channels and intermittent streams, sandy stream banks, open slopes, rocky shelves near seeps.	Moist open areas on gentle slopes in desert chaparral and pine transition areas.
Elevation	700–1300 m. (2300–4300 ft.)	1400–2700 m. (4600–8900 ft.)
Distribution	OR [WildflowerSearch map]	CA [WildflowerSearch map]
Discussion	Erythranthe washingtonensis is considered to be extirpated in Washington by the Washington Natural Heritage Program. (Discussion copyrighted by Flora of North America; reprinted with permission.)	Erythranthe discolor has two distinctive floral morphs; one is yellow with red spots on the palate, the other is pink with two yellow ridges on the palate. Populations can be monomorphic (usually yellow) or mixed, with the yellow morph most often in higher frequency. Erythranthe discolor was placed previously in synonymy with E. montioides, and the pink form of E. discolor has commonly been confused with E. palmeri. Erythranthe discolor is known to hybridize with E. barbata and is restricted to Kern and Tulare counties in the southern Sierra Nevada. (Discussion copyrighted by Flora of North America; reprinted with permission.)
Source	FNA vol. 17, p. 396.	FNA vol. 17, p. 386.
Parent taxa	Phrymaceae > Erythranthe	Phrymaceae > Erythranthe
Sibling taxa	E. acutidens, E. alsinoides, E. ampliata, E. androsacea, E. arenaria, E. arenicola, E. arvensis, E. barbata, E. bicolor, E. brachystylis, E. breviflora, E. breweri, E. caespitosa, E. calcicola, E. calciphila, E. cardinalis, E. carsonensis, E. charlestonensis, E. chinatiensis, E. cinnabarina, E. corallina, E. cordata, E. decora, E. dentata, E. diffusa, E. discolor, E. eastwoodiae, E. erubescens, E. exigua, E. filicaulis, E. filicifolia, E. floribunda, E. gemmipara, E. geniculata, E. geyeri, E. glaucescens, E. gracilipes, E. grandis, E. grayi, E. guttata, E. hallii, E. hardhamiae, E. hymenophylla, E. inamoena, E. inconspicua, E. inflatula, E. jungermannioides, E. laciniata, E. latidens, E. lewisii, E. linearifolia, E. marmorata, E. michiganensis, E. microphylla, E. minor, E. montioides, E. moschata, E. nasuta, E. norrisii, E. nudata, E. palmeri, E. pardalis, E. parishii, E. parvula, E. patula, E. percaulis, E. primuloides, E. ptilota, E. pulsiferae, E. purpurea, E. regni, E. rhodopetra, E. rubella, E. scouleri, E. shevockii, E. sierrae, E. suksdorfii, E. taylorii, E. thermalis, E. tilingii, E. trinitiensis, E. unimaculata, E. utahensis, E. verbenacea, E. willisii	E. acutidens, E. alsinoides, E. ampliata, E. androsacea, E. arenaria, E. arenicola, E. arvensis, E. barbata, E. bicolor, E. brachystylis, E. breviflora, E. breweri, E. caespitosa, E. calcicola, E. calciphila, E. cardinalis, E. carsonensis, E. charlestonensis, E. chinatiensis, E. cinnabarina, E. corallina, E. cordata, E. decora, E. dentata, E. diffusa, E. eastwoodiae, E. erubescens, E. exigua, E. filicaulis, E. filicifolia, E. floribunda, E. gemmipara, E. geniculata, E. geyeri, E. glaucescens, E. gracilipes, E. grandis, E. grayi, E. guttata, E. hallii, E. hardhamiae, E. hymenophylla, E. inamoena, E. inconspicua, E. inflatula, E. jungermannioides, E. laciniata, E. latidens, E. lewisii, E. linearifolia, E. marmorata, E. michiganensis, E. microphylla, E. minor, E. montioides, E. moschata, E. nasuta, E. norrisii, E. nudata, E. palmeri, E. pardalis, E. parishii, E. parvula, E. patula, E. percaulis, E. primuloides, E. ptilota, E. pulsiferae, E. purpurea, E. regni, E. rhodopetra, E. rubella, E. scouleri, E. shevockii, E. sierrae, E. suksdorfii, E. taylorii, E. thermalis, E. tilingii, E. trinitiensis, E. unimaculata, E. utahensis, E. verbenacea, E. washingtonensis, E. willisii
Synonyms	Mimulus washingtonensis	Mimulus discolor
Name authority	(Gandoger) G. L. Nesom: Phytoneuron 2012-39: 39. (2012) — (as washingtoniensis)	(A. L. Grant) N. S. Fraga: Phytoneuron 2012-39: 35. (2012)
Web links	Local floras: OR, WA Local Web sites: Flora NW, PNW Herbaria WildflowerSearch iNaturalist (observations) LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search	Local floras: CA Local Web sites: CalFlora, CalPhotos WildflowerSearch iNaturalist (observations) LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search

Erythranthe discolor

John Day or Washington monkeyflower, Washington monkey-flower

party-color monkeyflower, two-color monkeyflower

Habit

Annuals, fibrous-rooted or filiform-taprooted.

Annuals, taprooted.

Stems

erect to ascending, straight or geniculate at nodes, usually many-branched, terete, 5–25 cm, moderately puberulent-glandular to villous-glandular, hairs 0.1–0.8 mm, flattened, sometimes vitreous, distinctly multicellular, gland-tipped.

erect, simple or branched from basal nodes, 5–12 cm, sparsely glandular-pubescent, internodes elongate, distinct.

Leaves

cauline, basal not persistent;

petiole 2–14 mm;

blade palmately veined, deltate or ovate to ovate-lanceolate, 4–16(–23) × 2–11(–16) mm, base rounded to cuneate or truncate, margins denticulate or entire, apex acute, surfaces moderately puberulent-glandular to villous-glandular, hairs 0.1–0.8 mm, flattened, sometimes vitreous, distinctly multicellular, gland-tipped.

cauline, basal not persistent;

petiole 0–2 mm;

blade 1-veined or palmately 3-veined (in broader ones), linear to lanceolate or ovate, 5–15 × 1–4 mm, base attenuate, margins entire, apex acute, surfaces sparsely glandular-pubescent.

Flowers

herkogamous, 1–6, from proximal to distal nodes.

herkogamous, 1–8, from distal or medial to distal nodes.

Styles

hispid-hirtellous.

glabrous.

Corollas

yellow with small reddish brown dots, abaxial limb with 2 white patches (abaxial ridges), bilaterally symmetric, strongly bilabiate;

tube-throat funnelform, 8–10 mm, exserted beyond calyx margin;

limb expanded 7–10 mm, lobes obovate-oblong, apex rounded to rounded-cuneate.

yellow and tube-throat adaxial surface red-tinged, abaxial limb red-spotted or deep pink to purple and abaxial limb with 2 yellow ridges, bilaterally symmetric, strongly bilabiate;

tube-throat funnelform, 9–13 mm, exserted beyond calyx margin;

limb expanded 7–15 mm, lobes notched, adaxial limb glabrous or sparsely bearded.

Fruiting pedicels

divergent at nearly right angles, 20–50 mm, densely, minutely stipitate-glandular.

erect to ascending, 6–11(–32) mm.

Fruiting calyces

greenish, ridge-angled, tubular, weakly inflated, 6–8 mm, margins distinctly toothed or lobed, densely, minutely stipitate-glandular, lobes pronounced, erect.

sometimes red-dotted on ribs, campanulate, 4–8 mm, margins distinctly toothed or lobed, sparsely glandular-pubescent, ribs weak, lobes pronounced, erect, margins glabrous.

Capsules

included, 5–8.5 mm.

included, 5–8 mm.

Anthers

included, glabrous.

= 32.

Erythranthe washingtonensis

Erythranthe discolor

Phenology

Flowering May–Sep.

Flowering May–Jun.

Habitat

Shallow basalt gravel in narrow channels and intermittent streams, sandy stream banks, open slopes, rocky shelves near seeps.

Moist open areas on gentle slopes in desert chaparral and pine transition areas.

Elevation

700–1300 m. (2300–4300 ft.)

1400–2700 m. (4600–8900 ft.)

Distribution

[WildflowerSearch map]

Discussion

Erythranthe washingtonensis is considered to be extirpated in Washington by the Washington Natural Heritage Program.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Erythranthe discolor has two distinctive floral morphs; one is yellow with red spots on the palate, the other is pink with two yellow ridges on the palate. Populations can be monomorphic (usually yellow) or mixed, with the yellow morph most often in higher frequency.

Erythranthe discolor was placed previously in synonymy with E. montioides, and the pink form of E. discolor has commonly been confused with E. palmeri. Erythranthe discolor is known to hybridize with E. barbata and is restricted to Kern and Tulare counties in the southern Sierra Nevada.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source

FNA vol. 17, p. 396.

FNA vol. 17, p. 386.

Parent taxa

Phrymaceae > Erythranthe

Sibling taxa

E. acutidens, E. alsinoides, E. ampliata, E. androsacea, E. arenaria, E. arenicola, E. arvensis, E. barbata, E. bicolor, E. brachystylis, E. breviflora, E. breweri, E. caespitosa, E. calcicola, E. calciphila, E. cardinalis, E. carsonensis, E. charlestonensis, E. chinatiensis, E. cinnabarina, E. corallina, E. cordata, E. decora, E. dentata, E. diffusa, E. discolor, E. eastwoodiae, E. erubescens, E. exigua, E. filicaulis, E. filicifolia, E. floribunda, E. gemmipara, E. geniculata, E. geyeri, E. glaucescens, E. gracilipes, E. grandis, E. grayi, E. guttata, E. hallii, E. hardhamiae, E. hymenophylla, E. inamoena, E. inconspicua, E. inflatula, E. jungermannioides, E. laciniata, E. latidens, E. lewisii, E. linearifolia, E. marmorata, E. michiganensis, E. microphylla, E. minor, E. montioides, E. moschata, E. nasuta, E. norrisii, E. nudata, E. palmeri, E. pardalis, E. parishii, E. parvula, E. patula, E. percaulis, E. primuloides, E. ptilota, E. pulsiferae, E. purpurea, E. regni, E. rhodopetra, E. rubella, E. scouleri, E. shevockii, E. sierrae, E. suksdorfii, E. taylorii, E. thermalis, E. tilingii, E. trinitiensis, E. unimaculata, E. utahensis, E. verbenacea, E. willisii

E. acutidens, E. alsinoides, E. ampliata, E. androsacea, E. arenaria, E. arenicola, E. arvensis, E. barbata, E. bicolor, E. brachystylis, E. breviflora, E. breweri, E. caespitosa, E. calcicola, E. calciphila, E. cardinalis, E. carsonensis, E. charlestonensis, E. chinatiensis, E. cinnabarina, E. corallina, E. cordata, E. decora, E. dentata, E. diffusa, E. eastwoodiae, E. erubescens, E. exigua, E. filicaulis, E. filicifolia, E. floribunda, E. gemmipara, E. geniculata, E. geyeri, E. glaucescens, E. gracilipes, E. grandis, E. grayi, E. guttata, E. hallii, E. hardhamiae, E. hymenophylla, E. inamoena, E. inconspicua, E. inflatula, E. jungermannioides, E. laciniata, E. latidens, E. lewisii, E. linearifolia, E. marmorata, E. michiganensis, E. microphylla, E. minor, E. montioides, E. moschata, E. nasuta, E. norrisii, E. nudata, E. palmeri, E. pardalis, E. parishii, E. parvula, E. patula, E. percaulis, E. primuloides, E. ptilota, E. pulsiferae, E. purpurea, E. regni, E. rhodopetra, E. rubella, E. scouleri, E. shevockii, E. sierrae, E. suksdorfii, E. taylorii, E. thermalis, E. tilingii, E. trinitiensis, E. unimaculata, E. utahensis, E. verbenacea, E. washingtonensis, E. willisii

Synonyms

Mimulus washingtonensis

Mimulus discolor

Name authority

(Gandoger) G. L. Nesom: Phytoneuron 2012-39: 39. (2012) — (as washingtoniensis)

(A. L. Grant) N. S. Fraga: Phytoneuron 2012-39: 35. (2012)

Web links

Unless otherwise noted, all data comes from the online version of the Flora of North America, who have made most of their content available via a Creative Commons license (CC BY 4.0).

Choose a data source: Flora of North America, OregonFlora, UW Burke Herbarium

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Erythranthe washingtonensis

Erythranthe discolor

Erythranthe washingtonensis

Erythranthe discolor