FNA: Erythranthe washingtonensis vs. Erythranthe diffusa

	Erythranthe washingtonensis	Erythranthe diffusa
	John Day or Washington monkeyflower, Washington monkey-flower	Palomar monkeyflower
Habit	Annuals, fibrous-rooted or filiform-taprooted.	Annuals, taprooted.
Stems	erect to ascending, straight or geniculate at nodes, usually many-branched, terete, 5–25 cm, moderately puberulent-glandular to villous-glandular, hairs 0.1–0.8 mm, flattened, sometimes vitreous, distinctly multicellular, gland-tipped.	erect, simple or branched from basal nodes, 4–20 cm, minutely puberulent.
Leaves	cauline, basal not persistent; petiole 2–14 mm; blade palmately veined, deltate or ovate to ovate-lanceolate, 4–16(–23) × 2–11(–16) mm, base rounded to cuneate or truncate, margins denticulate or entire, apex acute, surfaces moderately puberulent-glandular to villous-glandular, hairs 0.1–0.8 mm, flattened, sometimes vitreous, distinctly multicellular, gland-tipped.	cauline, basal not persistent; petiole 0 mm; blade 1-veined or palmately 3-veined (in broader ones), lanceolate to ovate, 3–20 × 1–6 mm, base attenuate, margins entire, sometimes toothed, apex acute, surfaces minutely puberulent.
Flowers	herkogamous, 1–6, from proximal to distal nodes.	herkogamous, 1–36, from distal or medial to distal nodes.
Styles	hispid-hirtellous.	distally pubescent.
Corollas	yellow with small reddish brown dots, abaxial limb with 2 white patches (abaxial ridges), bilaterally symmetric, strongly bilabiate; tube-throat funnelform, 8–10 mm, exserted beyond calyx margin; limb expanded 7–10 mm, lobes obovate-oblong, apex rounded to rounded-cuneate.	pink to purple, abaxial limb with 2 yellow palate ridges, bilaterally symmetric, weakly bilabiate; tube-throat funnelform, (3–)6–10 mm, exserted beyond calyx margin; limb expanded 3–14 mm, lobes notched, adaxial limb glabrous, sometimes sparsely bearded.
Fruiting pedicels	divergent at nearly right angles, 20–50 mm, densely, minutely stipitate-glandular.	ascending to spreading horizontally, (2–)12–60(–68) mm.
Fruiting calyces	greenish, ridge-angled, tubular, weakly inflated, 6–8 mm, margins distinctly toothed or lobed, densely, minutely stipitate-glandular, lobes pronounced, erect.	sometimes red-dotted, campanulate, 3–6 mm, margins distinctly toothed or lobed, minutely puberulent, ribs weak, lobes pronounced, erect, margins glabrous.
Capsules	included, 5–8.5 mm.	included, 5–8 mm.
Anthers	included, glabrous.	included, glabrous.
Stigmas		distinctly shorter than corolla tube.
2n	= 32.

	Erythranthe washingtonensis	Erythranthe diffusa
Phenology	Flowering May–Sep.	Flowering Apr–Jun.
Habitat	Shallow basalt gravel in narrow channels and intermittent streams, sandy stream banks, open slopes, rocky shelves near seeps.	Moist areas in openings of chaparral, dry meadows in pine and oak woodlands.
Elevation	700–1300 m. (2300–4300 ft.)	300–1800 m. (1000–5900 ft.)
Distribution	OR [WildflowerSearch map]	CA; Mexico (Baja California) [WildflowerSearch map]
Discussion	Erythranthe washingtonensis is considered to be extirpated in Washington by the Washington Natural Heritage Program. (Discussion copyrighted by Flora of North America; reprinted with permission.)	Erythranthe diffusa has been included previously in E. palmeri but can be distinguished morphologically by glabrous margins on the calyx and pubescence on the distal end of the style. It occurs in Orange, Riverside, and San Diego counties as well as close to the California border in Baja California. (Discussion copyrighted by Flora of North America; reprinted with permission.)
Source	FNA vol. 17, p. 396.	FNA vol. 17, p. 387.
Parent taxa	Phrymaceae > Erythranthe	Phrymaceae > Erythranthe
Sibling taxa	E. acutidens, E. alsinoides, E. ampliata, E. androsacea, E. arenaria, E. arenicola, E. arvensis, E. barbata, E. bicolor, E. brachystylis, E. breviflora, E. breweri, E. caespitosa, E. calcicola, E. calciphila, E. cardinalis, E. carsonensis, E. charlestonensis, E. chinatiensis, E. cinnabarina, E. corallina, E. cordata, E. decora, E. dentata, E. diffusa, E. discolor, E. eastwoodiae, E. erubescens, E. exigua, E. filicaulis, E. filicifolia, E. floribunda, E. gemmipara, E. geniculata, E. geyeri, E. glaucescens, E. gracilipes, E. grandis, E. grayi, E. guttata, E. hallii, E. hardhamiae, E. hymenophylla, E. inamoena, E. inconspicua, E. inflatula, E. jungermannioides, E. laciniata, E. latidens, E. lewisii, E. linearifolia, E. marmorata, E. michiganensis, E. microphylla, E. minor, E. montioides, E. moschata, E. nasuta, E. norrisii, E. nudata, E. palmeri, E. pardalis, E. parishii, E. parvula, E. patula, E. percaulis, E. primuloides, E. ptilota, E. pulsiferae, E. purpurea, E. regni, E. rhodopetra, E. rubella, E. scouleri, E. shevockii, E. sierrae, E. suksdorfii, E. taylorii, E. thermalis, E. tilingii, E. trinitiensis, E. unimaculata, E. utahensis, E. verbenacea, E. willisii	E. acutidens, E. alsinoides, E. ampliata, E. androsacea, E. arenaria, E. arenicola, E. arvensis, E. barbata, E. bicolor, E. brachystylis, E. breviflora, E. breweri, E. caespitosa, E. calcicola, E. calciphila, E. cardinalis, E. carsonensis, E. charlestonensis, E. chinatiensis, E. cinnabarina, E. corallina, E. cordata, E. decora, E. dentata, E. discolor, E. eastwoodiae, E. erubescens, E. exigua, E. filicaulis, E. filicifolia, E. floribunda, E. gemmipara, E. geniculata, E. geyeri, E. glaucescens, E. gracilipes, E. grandis, E. grayi, E. guttata, E. hallii, E. hardhamiae, E. hymenophylla, E. inamoena, E. inconspicua, E. inflatula, E. jungermannioides, E. laciniata, E. latidens, E. lewisii, E. linearifolia, E. marmorata, E. michiganensis, E. microphylla, E. minor, E. montioides, E. moschata, E. nasuta, E. norrisii, E. nudata, E. palmeri, E. pardalis, E. parishii, E. parvula, E. patula, E. percaulis, E. primuloides, E. ptilota, E. pulsiferae, E. purpurea, E. regni, E. rhodopetra, E. rubella, E. scouleri, E. shevockii, E. sierrae, E. suksdorfii, E. taylorii, E. thermalis, E. tilingii, E. trinitiensis, E. unimaculata, E. utahensis, E. verbenacea, E. washingtonensis, E. willisii
Synonyms	Mimulus washingtonensis	Mimulus diffusus, M. grantianus
Name authority	(Gandoger) G. L. Nesom: Phytoneuron 2012-39: 39. (2012) — (as washingtoniensis)	(A. L. Grant) N. S. Fraga: Phytoneuron 2012-39: 35. (2012)
Web links	Local floras: OR, WA Local Web sites: Flora NW, PNW Herbaria WildflowerSearch iNaturalist (observations) LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search	Local floras: CA Local Web sites: CalFlora, CalPhotos WildflowerSearch iNaturalist (observations) LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search

Erythranthe diffusa

John Day or Washington monkeyflower, Washington monkey-flower

Palomar monkeyflower

Habit

Annuals, fibrous-rooted or filiform-taprooted.

Annuals, taprooted.

Stems

erect to ascending, straight or geniculate at nodes, usually many-branched, terete, 5–25 cm, moderately puberulent-glandular to villous-glandular, hairs 0.1–0.8 mm, flattened, sometimes vitreous, distinctly multicellular, gland-tipped.

erect, simple or branched from basal nodes, 4–20 cm, minutely puberulent.

Leaves

cauline, basal not persistent;

petiole 2–14 mm;

blade palmately veined, deltate or ovate to ovate-lanceolate, 4–16(–23) × 2–11(–16) mm, base rounded to cuneate or truncate, margins denticulate or entire, apex acute, surfaces moderately puberulent-glandular to villous-glandular, hairs 0.1–0.8 mm, flattened, sometimes vitreous, distinctly multicellular, gland-tipped.

cauline, basal not persistent;

petiole 0 mm;

blade 1-veined or palmately 3-veined (in broader ones), lanceolate to ovate, 3–20 × 1–6 mm, base attenuate, margins entire, sometimes toothed, apex acute, surfaces minutely puberulent.

Flowers

herkogamous, 1–6, from proximal to distal nodes.

herkogamous, 1–36, from distal or medial to distal nodes.

Styles

hispid-hirtellous.

distally pubescent.

Corollas

yellow with small reddish brown dots, abaxial limb with 2 white patches (abaxial ridges), bilaterally symmetric, strongly bilabiate;

tube-throat funnelform, 8–10 mm, exserted beyond calyx margin;

limb expanded 7–10 mm, lobes obovate-oblong, apex rounded to rounded-cuneate.

pink to purple, abaxial limb with 2 yellow palate ridges, bilaterally symmetric, weakly bilabiate;

tube-throat funnelform, (3–)6–10 mm, exserted beyond calyx margin;

limb expanded 3–14 mm, lobes notched, adaxial limb glabrous, sometimes sparsely bearded.

Fruiting pedicels

divergent at nearly right angles, 20–50 mm, densely, minutely stipitate-glandular.

ascending to spreading horizontally, (2–)12–60(–68) mm.

Fruiting calyces

greenish, ridge-angled, tubular, weakly inflated, 6–8 mm, margins distinctly toothed or lobed, densely, minutely stipitate-glandular, lobes pronounced, erect.

sometimes red-dotted, campanulate, 3–6 mm, margins distinctly toothed or lobed, minutely puberulent, ribs weak, lobes pronounced, erect, margins glabrous.

Capsules

included, 5–8.5 mm.

included, 5–8 mm.

Anthers

included, glabrous.

Stigmas

distinctly shorter than corolla tube.

= 32.

Erythranthe washingtonensis

Erythranthe diffusa

Phenology

Flowering May–Sep.

Flowering Apr–Jun.

Habitat

Shallow basalt gravel in narrow channels and intermittent streams, sandy stream banks, open slopes, rocky shelves near seeps.

Moist areas in openings of chaparral, dry meadows in pine and oak woodlands.

Elevation

700–1300 m. (2300–4300 ft.)

300–1800 m. (1000–5900 ft.)

Distribution

[WildflowerSearch map]

CA; Mexico (Baja California)

[WildflowerSearch map]

Discussion

Erythranthe washingtonensis is considered to be extirpated in Washington by the Washington Natural Heritage Program.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Erythranthe diffusa has been included previously in E. palmeri but can be distinguished morphologically by glabrous margins on the calyx and pubescence on the distal end of the style. It occurs in Orange, Riverside, and San Diego counties as well as close to the California border in Baja California.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source

FNA vol. 17, p. 396.

FNA vol. 17, p. 387.

Parent taxa

Phrymaceae > Erythranthe

Sibling taxa

E. acutidens, E. alsinoides, E. ampliata, E. androsacea, E. arenaria, E. arenicola, E. arvensis, E. barbata, E. bicolor, E. brachystylis, E. breviflora, E. breweri, E. caespitosa, E. calcicola, E. calciphila, E. cardinalis, E. carsonensis, E. charlestonensis, E. chinatiensis, E. cinnabarina, E. corallina, E. cordata, E. decora, E. dentata, E. diffusa, E. discolor, E. eastwoodiae, E. erubescens, E. exigua, E. filicaulis, E. filicifolia, E. floribunda, E. gemmipara, E. geniculata, E. geyeri, E. glaucescens, E. gracilipes, E. grandis, E. grayi, E. guttata, E. hallii, E. hardhamiae, E. hymenophylla, E. inamoena, E. inconspicua, E. inflatula, E. jungermannioides, E. laciniata, E. latidens, E. lewisii, E. linearifolia, E. marmorata, E. michiganensis, E. microphylla, E. minor, E. montioides, E. moschata, E. nasuta, E. norrisii, E. nudata, E. palmeri, E. pardalis, E. parishii, E. parvula, E. patula, E. percaulis, E. primuloides, E. ptilota, E. pulsiferae, E. purpurea, E. regni, E. rhodopetra, E. rubella, E. scouleri, E. shevockii, E. sierrae, E. suksdorfii, E. taylorii, E. thermalis, E. tilingii, E. trinitiensis, E. unimaculata, E. utahensis, E. verbenacea, E. willisii

E. acutidens, E. alsinoides, E. ampliata, E. androsacea, E. arenaria, E. arenicola, E. arvensis, E. barbata, E. bicolor, E. brachystylis, E. breviflora, E. breweri, E. caespitosa, E. calcicola, E. calciphila, E. cardinalis, E. carsonensis, E. charlestonensis, E. chinatiensis, E. cinnabarina, E. corallina, E. cordata, E. decora, E. dentata, E. discolor, E. eastwoodiae, E. erubescens, E. exigua, E. filicaulis, E. filicifolia, E. floribunda, E. gemmipara, E. geniculata, E. geyeri, E. glaucescens, E. gracilipes, E. grandis, E. grayi, E. guttata, E. hallii, E. hardhamiae, E. hymenophylla, E. inamoena, E. inconspicua, E. inflatula, E. jungermannioides, E. laciniata, E. latidens, E. lewisii, E. linearifolia, E. marmorata, E. michiganensis, E. microphylla, E. minor, E. montioides, E. moschata, E. nasuta, E. norrisii, E. nudata, E. palmeri, E. pardalis, E. parishii, E. parvula, E. patula, E. percaulis, E. primuloides, E. ptilota, E. pulsiferae, E. purpurea, E. regni, E. rhodopetra, E. rubella, E. scouleri, E. shevockii, E. sierrae, E. suksdorfii, E. taylorii, E. thermalis, E. tilingii, E. trinitiensis, E. unimaculata, E. utahensis, E. verbenacea, E. washingtonensis, E. willisii

Synonyms

Mimulus washingtonensis

Mimulus diffusus, M. grantianus

Name authority

(Gandoger) G. L. Nesom: Phytoneuron 2012-39: 39. (2012) — (as washingtoniensis)

(A. L. Grant) N. S. Fraga: Phytoneuron 2012-39: 35. (2012)

Web links

Unless otherwise noted, all data comes from the online version of the Flora of North America, who have made most of their content available via a Creative Commons license (CC BY 4.0).

Choose a data source: Flora of North America, OregonFlora, UW Burke Herbarium

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Flora of North America species comparison

Erythranthe washingtonensis

Erythranthe diffusa

Erythranthe washingtonensis

Erythranthe diffusa