FNA: Erythranthe washingtonensis vs. Erythranthe carsonensis

	Erythranthe washingtonensis	Erythranthe carsonensis
	John Day or Washington monkeyflower, Washington monkey-flower	Carson Valley monkeyflower
Habit	Annuals, fibrous-rooted or filiform-taprooted.	Annuals, taprooted, densely compact.
Stems	erect to ascending, straight or geniculate at nodes, usually many-branched, terete, 5–25 cm, moderately puberulent-glandular to villous-glandular, hairs 0.1–0.8 mm, flattened, sometimes vitreous, distinctly multicellular, gland-tipped.	erect, simple or branched from basal nodes, 2–7(–8) cm, minutely glandular, internodes shortened, not evident.
Leaves	cauline, basal not persistent; petiole 2–14 mm; blade palmately veined, deltate or ovate to ovate-lanceolate, 4–16(–23) × 2–11(–16) mm, base rounded to cuneate or truncate, margins denticulate or entire, apex acute, surfaces moderately puberulent-glandular to villous-glandular, hairs 0.1–0.8 mm, flattened, sometimes vitreous, distinctly multicellular, gland-tipped.	cauline, basal not persistent; petiole 0 mm; blade palmately 3-veined (in broader ones), linear to spatulate, (3–)5–23 × 1–5 mm, base truncate to truncate-cordate, clasping, margins entire, apex acute to obtuse, surfaces minutely glandular.
Flowers	herkogamous, 1–6, from proximal to distal nodes.	herkogamous, 1–35, from distal or medial to distal nodes.
Styles	hispid-hirtellous.	glabrous.
Corollas	yellow with small reddish brown dots, abaxial limb with 2 white patches (abaxial ridges), bilaterally symmetric, strongly bilabiate; tube-throat funnelform, 8–10 mm, exserted beyond calyx margin; limb expanded 7–10 mm, lobes obovate-oblong, apex rounded to rounded-cuneate.	yellow, palate red-dotted and 1 large central spot, bilaterally symmetric, strongly bilabiate; tube-throat cylindric, distinct from abruptly expanding throat, (5–)7–11 mm, exserted beyond calyx margin; limb expanded 7–12(–15) mm, each lobe 2-fid, palate densely bearded.
Fruiting pedicels	divergent at nearly right angles, 20–50 mm, densely, minutely stipitate-glandular.	(3–)5–14 mm.
Fruiting calyces	greenish, ridge-angled, tubular, weakly inflated, 6–8 mm, margins distinctly toothed or lobed, densely, minutely stipitate-glandular, lobes pronounced, erect.	campanulate to widely urceolate, 4–7 mm, margins distinctly toothed or lobed, minutely glandular, ribs thickened, lobes pronounced, erect.
Capsules	included, 5–8.5 mm.	included, 3–6 mm.
Anthers	included, glabrous.	included, glabrous.
2n	= 32.

	Erythranthe washingtonensis	Erythranthe carsonensis
Phenology	Flowering May–Sep.	Flowering Mar–Jun.
Habitat	Shallow basalt gravel in narrow channels and intermittent streams, sandy stream banks, open slopes, rocky shelves near seeps.	Openings in sage brush/bitterbrush scrub in sand of decomposed granite.
Elevation	700–1300 m. (2300–4300 ft.)	1400–1800 m. (4600–5900 ft.)
Distribution	OR [WildflowerSearch map]	CA; NV
Discussion	Erythranthe washingtonensis is considered to be extirpated in Washington by the Washington Natural Heritage Program. (Discussion copyrighted by Flora of North America; reprinted with permission.)	Erythranthe carsonensis is restricted to the Carson Valley, Eagle Valley, and Washoe Valley region of Nevada and adjacent California, with one known disjunct occurrence about 58 km to the north in Nevada. The species was previously included in the broader concept of E. montioides but can be distinguished by its much branched and compact habit, linear to spatulate leaves with clasping bases, calyx with glabrous margins, and larger corolla with one large red spot in the center. Erythranthe carsonensis has been impacted by agriculture, urbanization, and other anthropogenic changes. (Discussion copyrighted by Flora of North America; reprinted with permission.)
Source	FNA vol. 17, p. 396.	FNA vol. 17, p. 384.
Parent taxa	Phrymaceae > Erythranthe	Phrymaceae > Erythranthe
Sibling taxa	E. acutidens, E. alsinoides, E. ampliata, E. androsacea, E. arenaria, E. arenicola, E. arvensis, E. barbata, E. bicolor, E. brachystylis, E. breviflora, E. breweri, E. caespitosa, E. calcicola, E. calciphila, E. cardinalis, E. carsonensis, E. charlestonensis, E. chinatiensis, E. cinnabarina, E. corallina, E. cordata, E. decora, E. dentata, E. diffusa, E. discolor, E. eastwoodiae, E. erubescens, E. exigua, E. filicaulis, E. filicifolia, E. floribunda, E. gemmipara, E. geniculata, E. geyeri, E. glaucescens, E. gracilipes, E. grandis, E. grayi, E. guttata, E. hallii, E. hardhamiae, E. hymenophylla, E. inamoena, E. inconspicua, E. inflatula, E. jungermannioides, E. laciniata, E. latidens, E. lewisii, E. linearifolia, E. marmorata, E. michiganensis, E. microphylla, E. minor, E. montioides, E. moschata, E. nasuta, E. norrisii, E. nudata, E. palmeri, E. pardalis, E. parishii, E. parvula, E. patula, E. percaulis, E. primuloides, E. ptilota, E. pulsiferae, E. purpurea, E. regni, E. rhodopetra, E. rubella, E. scouleri, E. shevockii, E. sierrae, E. suksdorfii, E. taylorii, E. thermalis, E. tilingii, E. trinitiensis, E. unimaculata, E. utahensis, E. verbenacea, E. willisii	E. acutidens, E. alsinoides, E. ampliata, E. androsacea, E. arenaria, E. arenicola, E. arvensis, E. barbata, E. bicolor, E. brachystylis, E. breviflora, E. breweri, E. caespitosa, E. calcicola, E. calciphila, E. cardinalis, E. charlestonensis, E. chinatiensis, E. cinnabarina, E. corallina, E. cordata, E. decora, E. dentata, E. diffusa, E. discolor, E. eastwoodiae, E. erubescens, E. exigua, E. filicaulis, E. filicifolia, E. floribunda, E. gemmipara, E. geniculata, E. geyeri, E. glaucescens, E. gracilipes, E. grandis, E. grayi, E. guttata, E. hallii, E. hardhamiae, E. hymenophylla, E. inamoena, E. inconspicua, E. inflatula, E. jungermannioides, E. laciniata, E. latidens, E. lewisii, E. linearifolia, E. marmorata, E. michiganensis, E. microphylla, E. minor, E. montioides, E. moschata, E. nasuta, E. norrisii, E. nudata, E. palmeri, E. pardalis, E. parishii, E. parvula, E. patula, E. percaulis, E. primuloides, E. ptilota, E. pulsiferae, E. purpurea, E. regni, E. rhodopetra, E. rubella, E. scouleri, E. shevockii, E. sierrae, E. suksdorfii, E. taylorii, E. thermalis, E. tilingii, E. trinitiensis, E. unimaculata, E. utahensis, E. verbenacea, E. washingtonensis, E. willisii
Synonyms	Mimulus washingtonensis	Mimulus rubellus var. latiflorus
Name authority	(Gandoger) G. L. Nesom: Phytoneuron 2012-39: 39. (2012) — (as washingtoniensis)	N. S. Fraga: Aliso 30: 59, figs. 17–21. (2012)
Web links	Local floras: OR, WA Local Web sites: Flora NW, PNW Herbaria WildflowerSearch iNaturalist (observations) LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search	Local floras: CA Local Web sites: CalFlora, CalPhotos WildflowerSearch iNaturalist (observations) USDA Plants Database LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search

Erythranthe carsonensis

John Day or Washington monkeyflower, Washington monkey-flower

Carson Valley monkeyflower

Habit

Annuals, fibrous-rooted or filiform-taprooted.

Annuals, taprooted, densely compact.

Stems

erect to ascending, straight or geniculate at nodes, usually many-branched, terete, 5–25 cm, moderately puberulent-glandular to villous-glandular, hairs 0.1–0.8 mm, flattened, sometimes vitreous, distinctly multicellular, gland-tipped.

erect, simple or branched from basal nodes, 2–7(–8) cm, minutely glandular, internodes shortened, not evident.

Leaves

cauline, basal not persistent;

petiole 2–14 mm;

blade palmately veined, deltate or ovate to ovate-lanceolate, 4–16(–23) × 2–11(–16) mm, base rounded to cuneate or truncate, margins denticulate or entire, apex acute, surfaces moderately puberulent-glandular to villous-glandular, hairs 0.1–0.8 mm, flattened, sometimes vitreous, distinctly multicellular, gland-tipped.

cauline, basal not persistent;

petiole 0 mm;

blade palmately 3-veined (in broader ones), linear to spatulate, (3–)5–23 × 1–5 mm, base truncate to truncate-cordate, clasping, margins entire, apex acute to obtuse, surfaces minutely glandular.

Flowers

herkogamous, 1–6, from proximal to distal nodes.

herkogamous, 1–35, from distal or medial to distal nodes.

Styles

hispid-hirtellous.

glabrous.

Corollas

yellow with small reddish brown dots, abaxial limb with 2 white patches (abaxial ridges), bilaterally symmetric, strongly bilabiate;

tube-throat funnelform, 8–10 mm, exserted beyond calyx margin;

limb expanded 7–10 mm, lobes obovate-oblong, apex rounded to rounded-cuneate.

yellow, palate red-dotted and 1 large central spot, bilaterally symmetric, strongly bilabiate;

tube-throat cylindric, distinct from abruptly expanding throat, (5–)7–11 mm, exserted beyond calyx margin;

limb expanded 7–12(–15) mm, each lobe 2-fid, palate densely bearded.

Fruiting pedicels

divergent at nearly right angles, 20–50 mm, densely, minutely stipitate-glandular.

(3–)5–14 mm.

Fruiting calyces

greenish, ridge-angled, tubular, weakly inflated, 6–8 mm, margins distinctly toothed or lobed, densely, minutely stipitate-glandular, lobes pronounced, erect.

campanulate to widely urceolate, 4–7 mm, margins distinctly toothed or lobed, minutely glandular, ribs thickened, lobes pronounced, erect.

Capsules

included, 5–8.5 mm.

included, 3–6 mm.

Anthers

included, glabrous.

= 32.

Erythranthe washingtonensis

Erythranthe carsonensis

Phenology

Flowering May–Sep.

Flowering Mar–Jun.

Habitat

Shallow basalt gravel in narrow channels and intermittent streams, sandy stream banks, open slopes, rocky shelves near seeps.

Openings in sage brush/bitterbrush scrub in sand of decomposed granite.

Elevation

700–1300 m. (2300–4300 ft.)

1400–1800 m. (4600–5900 ft.)

Distribution

[WildflowerSearch map]

CA; NV

Discussion

Erythranthe washingtonensis is considered to be extirpated in Washington by the Washington Natural Heritage Program.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Erythranthe carsonensis is restricted to the Carson Valley, Eagle Valley, and Washoe Valley region of Nevada and adjacent California, with one known disjunct occurrence about 58 km to the north in Nevada. The species was previously included in the broader concept of E. montioides but can be distinguished by its much branched and compact habit, linear to spatulate leaves with clasping bases, calyx with glabrous margins, and larger corolla with one large red spot in the center. Erythranthe carsonensis has been impacted by agriculture, urbanization, and other anthropogenic changes.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source

FNA vol. 17, p. 396.

FNA vol. 17, p. 384.

Parent taxa

Phrymaceae > Erythranthe

Sibling taxa

E. acutidens, E. alsinoides, E. ampliata, E. androsacea, E. arenaria, E. arenicola, E. arvensis, E. barbata, E. bicolor, E. brachystylis, E. breviflora, E. breweri, E. caespitosa, E. calcicola, E. calciphila, E. cardinalis, E. carsonensis, E. charlestonensis, E. chinatiensis, E. cinnabarina, E. corallina, E. cordata, E. decora, E. dentata, E. diffusa, E. discolor, E. eastwoodiae, E. erubescens, E. exigua, E. filicaulis, E. filicifolia, E. floribunda, E. gemmipara, E. geniculata, E. geyeri, E. glaucescens, E. gracilipes, E. grandis, E. grayi, E. guttata, E. hallii, E. hardhamiae, E. hymenophylla, E. inamoena, E. inconspicua, E. inflatula, E. jungermannioides, E. laciniata, E. latidens, E. lewisii, E. linearifolia, E. marmorata, E. michiganensis, E. microphylla, E. minor, E. montioides, E. moschata, E. nasuta, E. norrisii, E. nudata, E. palmeri, E. pardalis, E. parishii, E. parvula, E. patula, E. percaulis, E. primuloides, E. ptilota, E. pulsiferae, E. purpurea, E. regni, E. rhodopetra, E. rubella, E. scouleri, E. shevockii, E. sierrae, E. suksdorfii, E. taylorii, E. thermalis, E. tilingii, E. trinitiensis, E. unimaculata, E. utahensis, E. verbenacea, E. willisii

E. acutidens, E. alsinoides, E. ampliata, E. androsacea, E. arenaria, E. arenicola, E. arvensis, E. barbata, E. bicolor, E. brachystylis, E. breviflora, E. breweri, E. caespitosa, E. calcicola, E. calciphila, E. cardinalis, E. charlestonensis, E. chinatiensis, E. cinnabarina, E. corallina, E. cordata, E. decora, E. dentata, E. diffusa, E. discolor, E. eastwoodiae, E. erubescens, E. exigua, E. filicaulis, E. filicifolia, E. floribunda, E. gemmipara, E. geniculata, E. geyeri, E. glaucescens, E. gracilipes, E. grandis, E. grayi, E. guttata, E. hallii, E. hardhamiae, E. hymenophylla, E. inamoena, E. inconspicua, E. inflatula, E. jungermannioides, E. laciniata, E. latidens, E. lewisii, E. linearifolia, E. marmorata, E. michiganensis, E. microphylla, E. minor, E. montioides, E. moschata, E. nasuta, E. norrisii, E. nudata, E. palmeri, E. pardalis, E. parishii, E. parvula, E. patula, E. percaulis, E. primuloides, E. ptilota, E. pulsiferae, E. purpurea, E. regni, E. rhodopetra, E. rubella, E. scouleri, E. shevockii, E. sierrae, E. suksdorfii, E. taylorii, E. thermalis, E. tilingii, E. trinitiensis, E. unimaculata, E. utahensis, E. verbenacea, E. washingtonensis, E. willisii

Synonyms

Mimulus washingtonensis

Mimulus rubellus var. latiflorus

Name authority

(Gandoger) G. L. Nesom: Phytoneuron 2012-39: 39. (2012) — (as washingtoniensis)

N. S. Fraga: Aliso 30: 59, figs. 17–21. (2012)

Web links

Unless otherwise noted, all data comes from the online version of the Flora of North America, who have made most of their content available via a Creative Commons license (CC BY 4.0).

Choose a data source: Flora of North America, OregonFlora, UW Burke Herbarium

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Flora of North America species comparison

Erythranthe washingtonensis

Erythranthe carsonensis

Erythranthe washingtonensis

Erythranthe carsonensis