Erythranthe washingtonensis |
Erythranthe calciphila |
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John Day or Washington monkeyflower, Washington monkey-flower |
Mexican moss monkeyflower |
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Habit | Annuals, fibrous-rooted or filiform-taprooted. | Annuals, usually fibrous-rooted, rarely rooting at proximal nodes. |
Stems | erect to ascending, straight or geniculate at nodes, usually many-branched, terete, 5–25 cm, moderately puberulent-glandular to villous-glandular, hairs 0.1–0.8 mm, flattened, sometimes vitreous, distinctly multicellular, gland-tipped. |
erect, sometimes decumbent-ascending, branched, 4–30 cm, minutely stipitate-glandular, also delicately villosulous-glandular along whole length. |
Leaves | cauline, basal not persistent; petiole 2–14 mm; blade palmately veined, deltate or ovate to ovate-lanceolate, 4–16(–23) × 2–11(–16) mm, base rounded to cuneate or truncate, margins denticulate or entire, apex acute, surfaces moderately puberulent-glandular to villous-glandular, hairs 0.1–0.8 mm, flattened, sometimes vitreous, distinctly multicellular, gland-tipped. |
basal and cauline; petiole 1–3 mm, cauline blade slightly or hardly reduced in size from basal, becoming subsessile to sessile (1–3 pairs of cauline leaves); blade palmately 3(–5)-veined, orbicular-ovate to oblong-ovate, 7–28 × 5–22 mm, base truncate to shallowly cuneate, margins shallowly dentate to denticulate, teeth 3–6 per side, apex rounded to obtuse, surfaces villous, hairs thin-walled, vitreous, eglandular or minutely gland-tipped, usually also minutely stipitate-glandular. |
Flowers | herkogamous, 1–6, from proximal to distal nodes. |
plesiogamous, 1–6(–10), axillary at all nodes, chasmogamous. |
Styles | hispid-hirtellous. |
glabrous. |
Corollas | yellow with small reddish brown dots, abaxial limb with 2 white patches (abaxial ridges), bilaterally symmetric, strongly bilabiate; tube-throat funnelform, 8–10 mm, exserted beyond calyx margin; limb expanded 7–10 mm, lobes obovate-oblong, apex rounded to rounded-cuneate. |
light yellow, red-spotted, weakly bilaterally symmetric, ± bilabiate; tube-throat narrowly funnelform, 5–7 mm, exserted 1–2 mm beyond calyx margin; limb expanded 2–4 mm. |
Fruiting pedicels | divergent at nearly right angles, 20–50 mm, densely, minutely stipitate-glandular. |
15–30(–55) mm, minutely stipitate-glandular, sometimes minutely hirtellous and minutely stipitate-glandular, sometimes short glandular-villous. |
Fruiting calyces | greenish, ridge-angled, tubular, weakly inflated, 6–8 mm, margins distinctly toothed or lobed, densely, minutely stipitate-glandular, lobes pronounced, erect. |
nodding 90º at maturity, usually 3(–5)-veined, ovoid, inflated, sagittally compressed, 6–10 mm, minutely stipitate-glandular, throat closing, lobes usually 3 or 3 and 5 on same plant, if 5 then with 2 interpolated lobes much smaller than abaxial pair. |
Capsules | included, 5–8.5 mm. |
included, 2–4 mm. |
Anthers | included, glabrous. |
included, glabrous. |
2n | = 32. |
= 30, 32. |
Erythranthe washingtonensis |
Erythranthe calciphila |
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Phenology | Flowering May–Sep. | Flowering Mar–Sep(–Nov). |
Habitat | Shallow basalt gravel in narrow channels and intermittent streams, sandy stream banks, open slopes, rocky shelves near seeps. | Rocky knobs, moist boulders, wet rock faces, roadcuts, seepages, springs, with moss, usually in pine or pine-oak woods. |
Elevation | 700–1300 m. (2300–4300 ft.) | 1800–2500 m. (5900–8200 ft.) |
Distribution |
OR
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AZ; Mexico (Chihuahua, Durango, Sinaloa, Sonora) |
Discussion | Erythranthe washingtonensis is considered to be extirpated in Washington by the Washington Natural Heritage Program. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Erythranthe calciphila is recognized by its annual duration (fibrous-rooted), short, erect stems with few, even-sized leaves (the basal often persistent), delicate stipitate-glandular vestiture, three-lobed calyces that are relatively large in fruit, small corollas, and autogamous reproduction. Plants rarely root at proximal nodes. In Arizona, at the northernmost extremity of its range, E. calciphila occurs in the Chiricahua, Huachuca, and Mule mountains of Cochise County. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 396. | FNA vol. 17, p. 424. |
Parent taxa | Phrymaceae > Erythranthe | Phrymaceae > Erythranthe |
Sibling taxa | ||
Synonyms | Mimulus washingtonensis | Mimulus calciphilus, M. minutiflorus |
Name authority | (Gandoger) G. L. Nesom: Phytoneuron 2012-39: 39. (2012) — (as washingtoniensis) | (Gentry) G. L. Nesom: Phytoneuron 2012-39: 43. (2012) |
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