Erythranthe washingtonensis |
Erythranthe brachystylis |
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John Day or Washington monkeyflower, Washington monkey-flower |
short-pedicel monkeyflower |
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Habit | Annuals, fibrous-rooted or filiform-taprooted. | Annuals, fibrous-rooted, sometimes taprooted, apparently sometimes producing thin runners from basal nodes. |
Stems | erect to ascending, straight or geniculate at nodes, usually many-branched, terete, 5–25 cm, moderately puberulent-glandular to villous-glandular, hairs 0.1–0.8 mm, flattened, sometimes vitreous, distinctly multicellular, gland-tipped. |
erect, simple or branched from proximal to medial nodes, 4-angled, filiform to slightly thickened, not distinctly fistulose, 6–22 cm, glabrous. |
Leaves | cauline, basal not persistent; petiole 2–14 mm; blade palmately veined, deltate or ovate to ovate-lanceolate, 4–16(–23) × 2–11(–16) mm, base rounded to cuneate or truncate, margins denticulate or entire, apex acute, surfaces moderately puberulent-glandular to villous-glandular, hairs 0.1–0.8 mm, flattened, sometimes vitreous, distinctly multicellular, gland-tipped. |
basal and cauline; petiole: proximals 1–8 mm, distals 0 mm; blade palmately 3–5-veined, ovate to depressed-ovate or suborbicular, 10–40 × 6–25 mm, base truncate to subcordate, margins undulate, subentire, or weakly, irregularly dentate, apex rounded, surfaces: proximals and medials glabrous, distals villous, hairs thin-walled, flattened, vitreous and sharp-pointed, eglandular. |
Flowers | herkogamous, 1–6, from proximal to distal nodes. |
plesiogamous, 4–10, from medial to distal nodes, cleistogamous. |
Styles | hispid-hirtellous. |
glabrous. |
Corollas | yellow with small reddish brown dots, abaxial limb with 2 white patches (abaxial ridges), bilaterally symmetric, strongly bilabiate; tube-throat funnelform, 8–10 mm, exserted beyond calyx margin; limb expanded 7–10 mm, lobes obovate-oblong, apex rounded to rounded-cuneate. |
yellow, without red markings, weakly bilaterally or nearly radially symmetric, weakly bilabiate or nearly regular; tube-throat narrowly cylindric, 7–9 mm, exserted 0–1 mm beyond calyx margin; limb expanded 3 mm. |
Fruiting pedicels | divergent at nearly right angles, 20–50 mm, densely, minutely stipitate-glandular. |
5–10 mm in proximal axils, shorter than or equal to subtending leaves, 1–5 mm distally and flowers and fruits appearing sessile or subsessile, glabrous. |
Fruiting calyces | greenish, ridge-angled, tubular, weakly inflated, 6–8 mm, margins distinctly toothed or lobed, densely, minutely stipitate-glandular, lobes pronounced, erect. |
red-tinged to sparsely purple-dotted or not, broadly elliptic-ovoid, inflated, sagittally compressed, 10–13 mm, minutely hirtellous, throat not or slightly closing. |
Capsules | included, 5–8.5 mm. |
included, stipitate, 4–5 mm. |
Anthers | included, glabrous. |
included, glabrous. |
2n | = 32. |
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Erythranthe washingtonensis |
Erythranthe brachystylis |
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Phenology | Flowering May–Sep. | Flowering Jun–Aug. |
Habitat | Shallow basalt gravel in narrow channels and intermittent streams, sandy stream banks, open slopes, rocky shelves near seeps. | Around springs, steep slopes. |
Elevation | 700–1300 m. (2300–4300 ft.) | 2100 m. (6900 ft.) |
Distribution |
OR
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NV |
Discussion | Erythranthe washingtonensis is considered to be extirpated in Washington by the Washington Natural Heritage Program. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Erythranthe brachystylis is closely similar to E. arvensis. Plants of both are annual in duration and produce depressed-ovate leaves, the distal with vitreous-villous surfaces, and tiny corollas barely exserted from the calyx and probably cleistogamous. Vestiture of the distal leaves includes an admixture of eglandular sharp-pointed hairs, sometimes encountered in E. arvensis, though not typical, perhaps reflecting introgression from E. nasuta. The distinction of Erythranthe brachystylis from E. arvensis is primarily in its foreshortened pedicels and more inflated fruiting calyces. The fruiting calyces appear to be subsessile or on pedicels shorter or only equaling the subtending leaves. The difference is essentially qualitative but produces a distinctive aspect. Erythranthe brachystylis is known only from the type collection in Nye County, a region where E. arvensis has not been documented. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 396. | FNA vol. 17, p. 422. |
Parent taxa | Phrymaceae > Erythranthe | Phrymaceae > Erythranthe |
Sibling taxa | ||
Synonyms | Mimulus washingtonensis | Mimulus brachystylis |
Name authority | (Gandoger) G. L. Nesom: Phytoneuron 2012-39: 39. (2012) — (as washingtoniensis) | (Edwin) G. L. Nesom: Phytoneuron 2012-39: 43. (2012) |
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