Erythranthe washingtonensis |
Erythranthe arenicola |
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John Day or Washington monkeyflower, Washington monkey-flower |
beach monkeyflower |
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Habit | Annuals, fibrous-rooted or filiform-taprooted. | Annuals, fibrous-rooted or slender-taprooted, rarely rooting at nodes. |
Stems | erect to ascending, straight or geniculate at nodes, usually many-branched, terete, 5–25 cm, moderately puberulent-glandular to villous-glandular, hairs 0.1–0.8 mm, flattened, sometimes vitreous, distinctly multicellular, gland-tipped. |
erect, rarely prostrate to prostrate-ascending, few-branched, 3–17 cm, moderately villous-glandular, hairs gland-tipped, or mixed hirtellous and stipitate-glandular. |
Leaves | cauline, basal not persistent; petiole 2–14 mm; blade palmately veined, deltate or ovate to ovate-lanceolate, 4–16(–23) × 2–11(–16) mm, base rounded to cuneate or truncate, margins denticulate or entire, apex acute, surfaces moderately puberulent-glandular to villous-glandular, hairs 0.1–0.8 mm, flattened, sometimes vitreous, distinctly multicellular, gland-tipped. |
basal and cauline; petiole: basal 2–8 mm or mid and distals absent; blade palmately 3–5-veined, suborbicular to broadly ovate or depressed-ovate, 5–17 × 6–15 mm, base truncate or truncate-cuneate to subcordate, margins subentire or crenulate, apex rounded to obtuse, surfaces moderately villous-glandular, hairs gland-tipped, or mixed hirtellous and stipitate-glandular. |
Flowers | herkogamous, 1–6, from proximal to distal nodes. |
herkogamous, 1–6, at distal nodes, chasmogamous. |
Styles | hispid-hirtellous. |
hirtellous. |
Corollas | yellow with small reddish brown dots, abaxial limb with 2 white patches (abaxial ridges), bilaterally symmetric, strongly bilabiate; tube-throat funnelform, 8–10 mm, exserted beyond calyx margin; limb expanded 7–10 mm, lobes obovate-oblong, apex rounded to rounded-cuneate. |
yellow, red-dotted, bilaterally symmetric, bilabiate; tube-throat funnelform, 11–20 mm, exserted 4–8 mm beyond calyx margin; limb expanded 10–18 mm. |
Fruiting pedicels | divergent at nearly right angles, 20–50 mm, densely, minutely stipitate-glandular. |
9–17 mm, moderately villous-glandular, hairs gland-tipped, or mixed hirtellous and stipitate-glandular. |
Fruiting calyces | greenish, ridge-angled, tubular, weakly inflated, 6–8 mm, margins distinctly toothed or lobed, densely, minutely stipitate-glandular, lobes pronounced, erect. |
nodding, ovoid-campanulate, inflated, sagittally compressed, 9–16 mm, moderately villous-glandular, hairs gland-tipped, or mixed hirtellous and stipitate-glandular, throat closing. |
Capsules | included, 5–8.5 mm. |
included, 5–12 mm. |
Anthers | included, glabrous. |
included, glabrous. |
2n | = 32. |
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Erythranthe washingtonensis |
Erythranthe arenicola |
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Phenology | Flowering May–Sep. | Flowering Apr–Aug. |
Habitat | Shallow basalt gravel in narrow channels and intermittent streams, sandy stream banks, open slopes, rocky shelves near seeps. | Sandy beaches, especially in moist hollows among dunes, sea cliff bases, chaparral near beaches, mudstone outcrops. |
Elevation | 700–1300 m. (2300–4300 ft.) | 0–100 m. (0–300 ft.) |
Distribution |
OR
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CA |
Discussion | Erythranthe washingtonensis is considered to be extirpated in Washington by the Washington Natural Heritage Program. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
F. W. Pennell (1947, 1951) considered Erythranthe arenicola an endemic of Monterey County, but plants from adjacent San Luis Obispo and Santa Cruz counties also belong here. Most of the localities are at seaside, but some are more than a mile inland. Erythranthe arenicola is hypothesized here to be a derivative of E. guttata or E. grandis, retaining the herkogamous breeding system of its putative ancestor but reduced in size and duration. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 396. | FNA vol. 17, p. 412. |
Parent taxa | Phrymaceae > Erythranthe | Phrymaceae > Erythranthe |
Sibling taxa | ||
Synonyms | Mimulus washingtonensis | Mimulus guttatus subsp. arenicola |
Name authority | (Gandoger) G. L. Nesom: Phytoneuron 2012-39: 39. (2012) — (as washingtoniensis) | (Pennell) G. L. Nesom: Phytoneuron 2012-39: 43. (2012) |
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