Erythranthe verbenacea |
Erythranthe arenaria |
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crimson monkeyflower, pico Pajaro |
sand-loving monkeyflower |
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Habit | Perennials, rhizomatous. | Annuals, fibrous-rooted or filiform-taprooted. |
Stems | erect to decumbent, usually simple, weakly 4-angled, 20–60 cm, ± glandular-villous. |
erect to ascending, straight or geniculate at nodes, simple or branched, 5–20 cm, villous-glandular, hairs 0.2–0.8 mm, gland-tipped. |
Leaves | cauline; petiole 0 mm; blade palmately 3–5-veined, elliptic to obovate, rhombic-ovate, or broadly spatulate, 50–75 × 15–26(–30) mm, base subcordate, subclasping, margins coarsely serrate, sometimes only distally, apex acute to obtuse, surfaces ± glandular-villous. |
basal and cauline; petiole 0 mm or proximals 1–3(–5) mm; blade 1-veined or palmately 3-veined, elliptic to narrowly elliptic, ovate-elliptic, or ovate-lanceolate, 5–12(–17) × 3–7 mm, base rounded to cuneate-attenuate, margins entire or sparsely dentate to serrate, apex acuminate to acute or obtuse, surfaces villous-glandular, hairs 0.2–0.8 mm, gland-tipped. |
Flowers | herkogamous, 2–12, axillary at leafy medial to distal nodes. |
herkogamous, 1–22, from proximal to distal nodes. |
Styles | glabrous. |
glabrous. |
Corollas | crimson, often yellow-tinged, palate ridges dark red, bilaterally symmetric, strongly bilabiate; tube-throat tubular, 25–35 mm, exserted 13–25 mm beyond calyx margin; abaxial limb spreading, adaxial erect, lobe apex truncate, often emarginate, throat open, palate ridges densely short-villous. |
yellow, abaxial limb red-dotted, bilaterally symmetric, weakly bilabiate; tube-throat funnelform, 9–12(–14) mm, exserted beyond calyx margin; lobes broadly obovate, apex rounded. |
Fruiting pedicels | 45–90(–150) mm. |
divergent-arcuate, 10–23 mm, villous-glandular, hairs 0.2–0.8 mm, gland-tipped. |
Fruiting calyces | campanulate, weakly inflated, 20–28 mm, sparsely glandular-villosulous to stipitate-glandular, lobes triangular to ovate-triangular, apex linear-triangular. |
usually red-dotted, narrowly campanulate, not or weakly inflated, 5–7(–9) mm, margins distinctly toothed or lobed, villous-glandular, ribs angled, lobes pronounced, erect. |
Capsules | included, 15–22 mm. |
included, 4–7 mm. |
Anthers | exserted, white-villous, thecae reflexed 45º. |
included, glabrous. |
2n | = 16. |
= 32. |
Erythranthe verbenacea |
Erythranthe arenaria |
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Phenology | Flowering Jun–Sep. | Flowering May–Sep. |
Habitat | Stream edges and beds, flood plains, around seeps and springs, canyon bottoms, moist cliff crevices and ledges. | Sandy flats, bars, gullies, washes, trails, roadcuts, seasonal creek beds and drainages, rocky slopes, seepy loam, ditches, lake edges, meadows, openings in pine-fir and pine-oak woodlands. |
Elevation | 300–2600 m. (1000–8500 ft.) | (100–)500–2600(–2800) m. ((300–)1600–8500(–9200) ft.) |
Distribution |
AZ; UT; Mexico (Baja California, Chihuahua, Durango, Sinaloa, Sonora)
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CA
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Discussion | R. K. Vickery (1992) noted that yellow-flowered morphs of Erythranthe verbenacea occur in a population at Vasey’s Paradise in the Grand Canyon (Coconino County), 32 miles downstream from Lees Ferry. Populations of Erythranthe verbenacea in the Oak Creek Canyon area in southern Coconino County, Arizona, have leaves with a narrow, lateral, undulating, purple stripe across the mid lamina. The coloration is retained even in dried specimens. In Utah, Erythranthe verbenacea is known only from the Zion Canyon area. Molecular (P. M. Beardsley et al. 2003) and morphological (G. L. Nesom 2014b) data indicate that Erythranthe verbenacea is sister to E. eastwoodiae. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Erythranthe arenaria is known from a cluster of six counties of the central Sierra Nevada: Fresno, Madera, Mariposa, Merced, Tulare, and Tuolumne. Most plants of Erythranthe arenaria have relatively even-sized cauline leaves, all sessile to proximally subsessile. Plants in the Yosemite area with persistent basal leaves that are short-petiolate, ovate with a cuneate base, and relatively larger than the more distal cauline ones, and possibly related to E. arenaria, have been named M. floribundus var. subulatus. These might be construed as showing the influence of E. geniculata, but that species occurs only at the lower range of elevation of E. arenaria, while plants referable to Mimulus floribundus var. subulatus occur at least to 2300 m and also have the erect habit characteristic of E. arenaria. These variants should be investigated, especially in the Yosemite area where they appear to be relatively common, with the possibility that they indeed represent a distinct entity. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 394. | FNA vol. 17, p. 403. |
Parent taxa | Phrymaceae > Erythranthe | Phrymaceae > Erythranthe |
Sibling taxa | ||
Synonyms | Mimulus verbenaceus, M. cardinalis var. verbenaceus, M. lugens | Mimulus arenarius, M. floribundus var. subulatus, M. multiflorus, M. subulatus, M. trisulcatus |
Name authority | (Greene) G. L. Nesom & N. S. Fraga: Phytoneuron 2012-39: 37. (2012) | (A. L. Grant) G. L. Nesom: Phytoneuron 2012-39: 38. (2012) |
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