Erythranthe tilingii |
Erythranthe utahensis |
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larger mountain monkeyflower, mountain monkey-flower, Tiling's monkey-flower |
Utah monkeyflower |
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Habit | Perennials, rhizomatous, solitary to weakly colonial, rhizomes forming a mass, yellowish, branching, filiform. | Perennials, rhizomatous, rooting at nodes, rhizomes filiform. |
Stems | erect-ascending, usually freely branched, 2–35 cm, glabrous or sparsely stipitate-glandular to short glandular-villous. |
erect, sometimes decumbent-ascending proximally, simple or few-branched, 20–50 cm, glabrous or sparsely stipitate-glandular in inflorescence. |
Leaves | cauline; petiole 0–25 mm, distals 0 mm; blade palmately 3–5-veined, ovate to lanceolate-triangular or narrowly lanceolate (broadly ovate in large-leaved forms), 5–35(–55) mm, base cuneate to attenuate, margins irregularly denticulate, apex acute to obtuse or rounded, surfaces glabrous, sparsely stipitate-glandular to short glandular-villous, glabrate, or sparsely to moderately villous, hairs thick-vitreous, eglandular. |
basal and cauline, even-sized or largest near mid stem; petiole 0 mm or proximalmost 2–10 mm; blade palmately 3–5-veined, orbicular or suborbicular to broadly elliptic, broadly ovate, or depressed-ovate, 20–40(–75) × 12–35(–40) mm, base usually truncate to broadly cuneate, margins entire or subentire to mucronulate, shallowly dentate, or denticulate, apex rounded, surfaces glabrous or glabrate to sparsely stipitate-glandular and sparsely pilose, hairs thin-walled, abaxial often glaucous. |
Flowers | herkogamous, 1–3(–5), from distal nodes. |
herkogamous, 6–16, from proximal to distal nodes, in a loose raceme, distal bracts becoming much reduced. |
Styles | hirtellous. |
hirtellous. |
Corollas | yellow, red-dotted, bilaterally symmetric, bilabiate; tube-throat broadly funnelform, 15–28 mm, exserted 5–10 mm beyond calyx margin; limb expanded 14–30 mm. |
yellow, abaxial limb prominently darker yellow, sparsely purple-spotted, bilaterally symmetric, weakly bilabiate; tube-throat narrowly funnelform to broadly cylindric, 10–15 mm, exserted 5–8 mm beyond calyx margin; limb expanded 12–20 mm. |
Fruiting pedicels | 15–35(–40) mm, sparsely stipitate-glandular to short glandular-villous. |
(25–)40–75 mm, stipitate-glandular to short villous, hairs gland-tipped. |
Fruiting calyces | usually purple-tinged and purple-dotted, broadly campanulate, inflated, sagittally compressed, 11–15 mm, glabrous or sparsely stipitate-glandular to short glandular-villous, villous at sinuses, throat closing, lobes broadly ovate, abaxial usually longer than lateral, adaxial at least 2 times as long as others. |
broadly ovate-cylindric, inflated, sagittally compressed, (10–)11–17(–20) mm, stipitate-glandular or minutely hirtellous or a mixture, hairs sometimes also longer, thin-walled, eglandular or glandular, throat not closing, adaxial lobe slightly longer, triangular-blunt. |
Capsules | included, 5–7 mm. |
included, 5–7 mm. |
Anthers | included, glabrous. |
included, glabrous. |
2n | = 28, 56. |
= 28, 30. |
Erythranthe tilingii |
Erythranthe utahensis |
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Phenology | Flowering Jul–Sep. | Flowering (Feb–)May–Aug(–Oct). |
Habitat | Seeps, springs, stream banks, shallow rivulets, cliff bases, ledges and crevices, steep gravelly slopes, wet meadows. | Drainage ditches, springs, seeps, wet meadows, margins of ponds and small streams, marshy areas. |
Elevation | 1400–3400 m. (4600–11200 ft.) | 1400–2500 m. (4600–8200 ft.) |
Distribution |
AZ; CA; ID; MT; NV; OR; UT; AB
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CA; CO; NV; UT |
Discussion | Plants of Erythranthe tilingii are characterized by their relatively low stature and stems arising from a system of thin rhizomes and producing mostly one to three large flowers each; they usually occur at relatively high elevations. Erythranthe tilingii sometimes has been considered to include one or several infraspecific entities; from within this taxonomic amalgam, four distinct species are recognized here: E. caespitosa, E. corallina, E. minor, and E. tilingii. Erythranthe corallina and E. minor probably are more closely related to E. guttata. The populations identified here as E. tilingii from northeastern Oregon northeast to Alberta and southeast to Utah may prove to be a separate (undescribed) species. Erythranthe tilingii in the strict sense is relatively widespread over the western United States and is sympatric with E. caespitosa and E. corallina. Leaves in E. tilingii are variable in size, and particularly in Idaho, they may approach the small size of those of E. caespitosa, but the leaf margins of E. tilingii are distinctly toothed, and the stems are taller and more erect. Across the range of the species, plants sometimes produce very large leaves, but these often occur on plants with characteristically smaller leaves. This wide variability in size apparently does not occur in E. caespitosa. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Erythranthe utahensis is characterized by its erect stems, prolifically produced filiform rhizomes, basal leaves short-petiolate to subsessile and cauline sessile, blades suborbicular to broadly ovate or depressed ovate with thin-walled villous-glandular hairs on both surfaces, margins subentire, proximal pedicels elongating to 75 mm, and calyces open at maturity. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 408. | FNA vol. 17, p. 410. |
Parent taxa | Phrymaceae > Erythranthe | Phrymaceae > Erythranthe |
Sibling taxa | ||
Synonyms | Mimulus tilingii, M. caespitosus var. implexus, M. implexus, M. implicatus, M. langsdorffii var. tilingii, M. lucens, M. veronicifolius | Mimulus glabratus subsp. utahensis |
Name authority | (Regel) G. L. Nesom: Phytoneuron 2012-39: 44. (2012) | (Pennell) G. L. Nesom: Phytoneuron 2012-39: 44. (2012) |
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