Erythranthe tilingii |
Erythranthe sierrae |
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larger mountain monkeyflower, mountain monkey-flower, Tiling's monkey-flower |
Sierra Nevada monkeyflower |
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Habit | Perennials, rhizomatous, solitary to weakly colonial, rhizomes forming a mass, yellowish, branching, filiform. | Annuals, taprooted. |
Stems | erect-ascending, usually freely branched, 2–35 cm, glabrous or sparsely stipitate-glandular to short glandular-villous. |
erect, simple or branched from basal nodes, (3–)4–20 cm, minutely glandular. |
Leaves | cauline; petiole 0–25 mm, distals 0 mm; blade palmately 3–5-veined, ovate to lanceolate-triangular or narrowly lanceolate (broadly ovate in large-leaved forms), 5–35(–55) mm, base cuneate to attenuate, margins irregularly denticulate, apex acute to obtuse or rounded, surfaces glabrous, sparsely stipitate-glandular to short glandular-villous, glabrate, or sparsely to moderately villous, hairs thick-vitreous, eglandular. |
cauline, basal not persistent; petiole 0 mm; blade 1-veined or palmately 3-veined (in broader ones), linear to oblanceolate, 3–23 × 1–10 mm, base attenuate, margins entire, sometimes toothed, apex acute, surfaces minutely glandular. |
Flowers | herkogamous, 1–3(–5), from distal nodes. |
herkogamous, 1–38, from distal or medial to distal nodes. |
Styles | hirtellous. |
glabrous. |
Corollas | yellow, red-dotted, bilaterally symmetric, bilabiate; tube-throat broadly funnelform, 15–28 mm, exserted 5–10 mm beyond calyx margin; limb expanded 14–30 mm. |
pale pink, abaxial limb with 2 yellow ridges, bilaterally symmetric, strongly bilabiate; tube-throat funnelform, 5–15 mm, exserted beyond calyx margin; limb expanded 5–17 mm, lobes deeply notched, abaxial limb glabrous or sparsely puberulent. |
Fruiting pedicels | 15–35(–40) mm, sparsely stipitate-glandular to short glandular-villous. |
10–33(–40) mm. |
Fruiting calyces | usually purple-tinged and purple-dotted, broadly campanulate, inflated, sagittally compressed, 11–15 mm, glabrous or sparsely stipitate-glandular to short glandular-villous, villous at sinuses, throat closing, lobes broadly ovate, abaxial usually longer than lateral, adaxial at least 2 times as long as others. |
sometimes red-spotted, campanulate to cylindric, 4–8 mm, margins distinctly toothed or lobed, minutely glandular, ribs weak, lobes pronounced, erect, margins ciliate. |
Capsules | included, 5–7 mm. |
included, 4–9 mm. |
Anthers | included, glabrous. |
included, glabrous. |
2n | = 28, 56. |
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Erythranthe tilingii |
Erythranthe sierrae |
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Phenology | Flowering Jul–Sep. | Flowering Apr–Jul. |
Habitat | Seeps, springs, stream banks, shallow rivulets, cliff bases, ledges and crevices, steep gravelly slopes, wet meadows. | Foothill oak woodlands, mixed coniferous forests. |
Elevation | 1400–3400 m. (4600–11200 ft.) | 200–2300 m. (700–7500 ft.) |
Distribution |
AZ; CA; ID; MT; NV; OR; UT; AB
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CA |
Discussion | Plants of Erythranthe tilingii are characterized by their relatively low stature and stems arising from a system of thin rhizomes and producing mostly one to three large flowers each; they usually occur at relatively high elevations. Erythranthe tilingii sometimes has been considered to include one or several infraspecific entities; from within this taxonomic amalgam, four distinct species are recognized here: E. caespitosa, E. corallina, E. minor, and E. tilingii. Erythranthe corallina and E. minor probably are more closely related to E. guttata. The populations identified here as E. tilingii from northeastern Oregon northeast to Alberta and southeast to Utah may prove to be a separate (undescribed) species. Erythranthe tilingii in the strict sense is relatively widespread over the western United States and is sympatric with E. caespitosa and E. corallina. Leaves in E. tilingii are variable in size, and particularly in Idaho, they may approach the small size of those of E. caespitosa, but the leaf margins of E. tilingii are distinctly toothed, and the stems are taller and more erect. Across the range of the species, plants sometimes produce very large leaves, but these often occur on plants with characteristically smaller leaves. This wide variability in size apparently does not occur in E. caespitosa. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Erythranthe sierrae is endemic to the southern Sierra Nevada in Fresno, Kern, and Tulare counties. The species was previously treated as E. palmeri but can be distinguished by having leaf margins that are often toothed, pale pink corollas, and white stamens. Erythranthe palmeri has entire leaf margins that are never toothed, deep pink to purple corollas, and yellow stamens. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 408. | FNA vol. 17, p. 385. |
Parent taxa | Phrymaceae > Erythranthe | Phrymaceae > Erythranthe |
Sibling taxa | ||
Synonyms | Mimulus tilingii, M. caespitosus var. implexus, M. implexus, M. implicatus, M. langsdorffii var. tilingii, M. lucens, M. veronicifolius | |
Name authority | (Regel) G. L. Nesom: Phytoneuron 2012-39: 44. (2012) | N. S. Fraga: Aliso 30: 67, figs. 29–31. (2012) |
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