Erythranthe tilingii |
Erythranthe regni |
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larger mountain monkeyflower, mountain monkey-flower, Tiling's monkey-flower |
King of Arizona monkeyflower |
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Habit | Perennials, rhizomatous, solitary to weakly colonial, rhizomes forming a mass, yellowish, branching, filiform. | Annuals, fibrous-rooted, sometimes rooting at proximal nodes. |
Stems | erect-ascending, usually freely branched, 2–35 cm, glabrous or sparsely stipitate-glandular to short glandular-villous. |
erect to ascending-erect, branched, sometimes becoming slightly fistulose, 15–45 cm, glabrous. |
Leaves | cauline; petiole 0–25 mm, distals 0 mm; blade palmately 3–5-veined, ovate to lanceolate-triangular or narrowly lanceolate (broadly ovate in large-leaved forms), 5–35(–55) mm, base cuneate to attenuate, margins irregularly denticulate, apex acute to obtuse or rounded, surfaces glabrous, sparsely stipitate-glandular to short glandular-villous, glabrate, or sparsely to moderately villous, hairs thick-vitreous, eglandular. |
basal and cauline; petiole: proximals 5–25(–30) mm, mid cauline and distals not connate, 0 mm; blade palmately 5–7-veined, proximal sometimes subpinnate, proximals ovate to depressed-orbicular, 15–20(–50) × 15–25(–50) mm, medials and distals broadly depressed-ovate to obtriangular or flabellate, 15–35 mm, largest basal or at mid stem with distal slightly reduced, base attenuate-cuneate, margins shallowly serrate-dentate, sometimes irregularly, to mucronulate or apiculate, teeth (3–)5–7 per side, rarely subentire, apex rounded, surfaces glabrous. |
Flowers | herkogamous, 1–3(–5), from distal nodes. |
plesiogamous, 6–16, from all nodes or medial to distal, cleistogamous. |
Styles | hirtellous. |
glabrous. |
Corollas | yellow, red-dotted, bilaterally symmetric, bilabiate; tube-throat broadly funnelform, 15–28 mm, exserted 5–10 mm beyond calyx margin; limb expanded 14–30 mm. |
yellow, not red-dotted, bilaterally or radially symmetric, bilabiate or regular; tube-throat cylindric-funnelform, 9–12 mm, exserted 3–5 mm beyond calyx margin; limb expanded 1–1.5 mm. |
Fruiting pedicels | 15–35(–40) mm, sparsely stipitate-glandular to short glandular-villous. |
15–30 mm, longer than subtending leaves, glabrous. |
Fruiting calyces | usually purple-tinged and purple-dotted, broadly campanulate, inflated, sagittally compressed, 11–15 mm, glabrous or sparsely stipitate-glandular to short glandular-villous, villous at sinuses, throat closing, lobes broadly ovate, abaxial usually longer than lateral, adaxial at least 2 times as long as others. |
sparsely purple-dotted, broadly campanulate-cylindric, inflated, sagittally compressed, 7–9 mm, glabrous, throat not closing, adaxial lobe longest. |
Capsules | included, 5–7 mm. |
included, 4–5 mm. |
Anthers | included, glabrous. |
included, glabrous. |
2n | = 28, 56. |
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Erythranthe tilingii |
Erythranthe regni |
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Phenology | Flowering Jul–Sep. | Flowering Mar–May. |
Habitat | Seeps, springs, stream banks, shallow rivulets, cliff bases, ledges and crevices, steep gravelly slopes, wet meadows. | Moist to wet, sandy loam soils. |
Elevation | 1400–3400 m. (4600–11200 ft.) | 800–1000 m. (2600–3300 ft.) |
Distribution |
AZ; CA; ID; MT; NV; OR; UT; AB
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AZ |
Discussion | Plants of Erythranthe tilingii are characterized by their relatively low stature and stems arising from a system of thin rhizomes and producing mostly one to three large flowers each; they usually occur at relatively high elevations. Erythranthe tilingii sometimes has been considered to include one or several infraspecific entities; from within this taxonomic amalgam, four distinct species are recognized here: E. caespitosa, E. corallina, E. minor, and E. tilingii. Erythranthe corallina and E. minor probably are more closely related to E. guttata. The populations identified here as E. tilingii from northeastern Oregon northeast to Alberta and southeast to Utah may prove to be a separate (undescribed) species. Erythranthe tilingii in the strict sense is relatively widespread over the western United States and is sympatric with E. caespitosa and E. corallina. Leaves in E. tilingii are variable in size, and particularly in Idaho, they may approach the small size of those of E. caespitosa, but the leaf margins of E. tilingii are distinctly toothed, and the stems are taller and more erect. Across the range of the species, plants sometimes produce very large leaves, but these often occur on plants with characteristically smaller leaves. This wide variability in size apparently does not occur in E. caespitosa. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Erythranthe regni is endemic to the Kofa Mountains of Yuma County; all collections have been made from the Kofa Game Refuge (Kofa National Wildlife Refuge). Because its calyces remain open at maturity, this species is hypothesized to be most closely related to E. geyeri, from which it differs by its erect habit, apparently annual duration, larger leaves, purple-dotted calyces, and corollas with longer tube-throat and barely bilabiate limb. Geography and other morphology, however, suggest that its evolutionary origins are closer to E. guttata. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 408. | FNA vol. 17, p. 406. |
Parent taxa | Phrymaceae > Erythranthe | Phrymaceae > Erythranthe |
Sibling taxa | ||
Synonyms | Mimulus tilingii, M. caespitosus var. implexus, M. implexus, M. implicatus, M. langsdorffii var. tilingii, M. lucens, M. veronicifolius | |
Name authority | (Regel) G. L. Nesom: Phytoneuron 2012-39: 44. (2012) | G. L. Nesom: Phytoneuron 2012-40: 24. (2012) |
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