Erythranthe tilingii |
Erythranthe parvula |
|
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larger mountain monkeyflower, mountain monkey-flower, Tiling's monkey-flower |
Southwestern mat monkeyflower |
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Habit | Perennials, rhizomatous, solitary to weakly colonial, rhizomes forming a mass, yellowish, branching, filiform. | Perennials, rhizomatous, sometimes rooting at nodes, mat-forming. |
Stems | erect-ascending, usually freely branched, 2–35 cm, glabrous or sparsely stipitate-glandular to short glandular-villous. |
procumbent, branched, 5–15 cm, stipitate-glandular distally. |
Leaves | cauline; petiole 0–25 mm, distals 0 mm; blade palmately 3–5-veined, ovate to lanceolate-triangular or narrowly lanceolate (broadly ovate in large-leaved forms), 5–35(–55) mm, base cuneate to attenuate, margins irregularly denticulate, apex acute to obtuse or rounded, surfaces glabrous, sparsely stipitate-glandular to short glandular-villous, glabrate, or sparsely to moderately villous, hairs thick-vitreous, eglandular. |
cauline; petiole 1–4 mm; blade palmately 3-veined, ovate to orbicular-ovate or depressed-ovate, 3–11 × 3–9 mm, base truncate to cuneate, margins shallowly denticulate to dentate, teeth 3–5 per side, apex acute to obtuse, surfaces villous-hirsute, hairs whitish, thickened, flattened, stiff, gland-tipped. |
Flowers | herkogamous, 1–3(–5), from distal nodes. |
plesiogamous, 2–8, axillary at distal nodes. |
Styles | hirtellous. |
glabrous. |
Corollas | yellow, red-dotted, bilaterally symmetric, bilabiate; tube-throat broadly funnelform, 15–28 mm, exserted 5–10 mm beyond calyx margin; limb expanded 14–30 mm. |
yellow, red-dotted, bilaterally symmetric, bilabiate; tube-throat funnelform, 6–8 mm, exserted beyond calyx margin; limb expanded 4–6 mm, abaxial limb spreading, lobes fimbriate. |
Fruiting pedicels | 15–35(–40) mm, sparsely stipitate-glandular to short glandular-villous. |
7–15 mm, minutely stipitate-glandular. |
Fruiting calyces | usually purple-tinged and purple-dotted, broadly campanulate, inflated, sagittally compressed, 11–15 mm, glabrous or sparsely stipitate-glandular to short glandular-villous, villous at sinuses, throat closing, lobes broadly ovate, abaxial usually longer than lateral, adaxial at least 2 times as long as others. |
nodding 45–90º, 5-lobed, cylindric-ovoid, inflated, sagittally compressed, 4–5 mm, villous-hirsute, throat closing. |
Capsules | included, 5–7 mm. |
included, 3–4 mm. |
Anthers | included, glabrous. |
included, glabrous. |
2n | = 28, 56. |
= 32. |
Erythranthe tilingii |
Erythranthe parvula |
|
Phenology | Flowering Jul–Sep. | Flowering Apr–Sep. |
Habitat | Seeps, springs, stream banks, shallow rivulets, cliff bases, ledges and crevices, steep gravelly slopes, wet meadows. | Wet vertical rock faces, ledges, and rocky slopes, seepy wash banks. |
Elevation | 1400–3400 m. (4600–11200 ft.) | 500–2400(–3400) m. (1600–7900(–11200) ft.) |
Distribution |
AZ; CA; ID; MT; NV; OR; UT; AB
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AZ; NM; Mexico (Sonora) |
Discussion | Plants of Erythranthe tilingii are characterized by their relatively low stature and stems arising from a system of thin rhizomes and producing mostly one to three large flowers each; they usually occur at relatively high elevations. Erythranthe tilingii sometimes has been considered to include one or several infraspecific entities; from within this taxonomic amalgam, four distinct species are recognized here: E. caespitosa, E. corallina, E. minor, and E. tilingii. Erythranthe corallina and E. minor probably are more closely related to E. guttata. The populations identified here as E. tilingii from northeastern Oregon northeast to Alberta and southeast to Utah may prove to be a separate (undescribed) species. Erythranthe tilingii in the strict sense is relatively widespread over the western United States and is sympatric with E. caespitosa and E. corallina. Leaves in E. tilingii are variable in size, and particularly in Idaho, they may approach the small size of those of E. caespitosa, but the leaf margins of E. tilingii are distinctly toothed, and the stems are taller and more erect. Across the range of the species, plants sometimes produce very large leaves, but these often occur on plants with characteristically smaller leaves. This wide variability in size apparently does not occur in E. caespitosa. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
All monkeyflowers with laciniate-lobed corollas have generally been identified as Mimulus dentilobus B. L. Robinson & Fernald, but these occur as three, morphologically distinct, widely allopatric population systems, each of which is regarded as a separate species (G. L. Nesom 2012g). Erythranthe dentiloba (B. L. Robinson & Fernald) G. L. Nesom, which is endemic to Mexico, is the only one of the three with an allogamous breeding system. Erythranthe parvula is restricted to Arizona and New Mexico except for one locality in northern Sonora, Mexico. Erythranthe chinatiensis is the third species of the group. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 408. | FNA vol. 17, p. 424. |
Parent taxa | Phrymaceae > Erythranthe | Phrymaceae > Erythranthe |
Sibling taxa | ||
Synonyms | Mimulus tilingii, M. caespitosus var. implexus, M. implexus, M. implicatus, M. langsdorffii var. tilingii, M. lucens, M. veronicifolius | Mimulus parvulus |
Name authority | (Regel) G. L. Nesom: Phytoneuron 2012-39: 44. (2012) | (Wooton & Standley) G. L. Nesom: Phytoneuron 2012-39: 44. (2012) |
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