Erythranthe tilingii |
Erythranthe hymenophylla |
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larger mountain monkeyflower, mountain monkey-flower, Tiling's monkey-flower |
thin-sepal monkeyflower |
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Habit | Perennials, rhizomatous, solitary to weakly colonial, rhizomes forming a mass, yellowish, branching, filiform. | Annuals, filiform-taprooted. |
Stems | erect-ascending, usually freely branched, 2–35 cm, glabrous or sparsely stipitate-glandular to short glandular-villous. |
prostrate to ascending-erect, sharply bent at basal nodes, simple or few-branched, 5–25 cm, glandular-puberulent to glandular-villous, hairs 0.1–0.8 mm, vitreous, flattened, multicellular, gland-tipped. |
Leaves | cauline; petiole 0–25 mm, distals 0 mm; blade palmately 3–5-veined, ovate to lanceolate-triangular or narrowly lanceolate (broadly ovate in large-leaved forms), 5–35(–55) mm, base cuneate to attenuate, margins irregularly denticulate, apex acute to obtuse or rounded, surfaces glabrous, sparsely stipitate-glandular to short glandular-villous, glabrate, or sparsely to moderately villous, hairs thick-vitreous, eglandular. |
basal and cauline, largest at mid stem; petiole 6–30 mm; blade pinnately veined, broadly lanceolate to ovate, 10–35 × 10–30 mm, distinctly membranous, base cuneate to shallowly cordate, margins coarsely dentate to shallowly denticulate or entire, apex acute to obtuse, surfaces glandular-puberulent to glandular-villous, hairs 0.1–0.8 mm, vitreous, flattened, multicellular, gland-tipped, glandular. |
Flowers | herkogamous, 1–3(–5), from distal nodes. |
herkogamous, 1–6, from proximal to distal nodes. |
Styles | hirtellous. |
glabrous. |
Corollas | yellow, red-dotted, bilaterally symmetric, bilabiate; tube-throat broadly funnelform, 15–28 mm, exserted 5–10 mm beyond calyx margin; limb expanded 14–30 mm. |
light yellow, throat and abaxial lobes red- or purple-spotted, sometimes with small white patches, bilaterally symmetric, weakly bilabiate; tube-throat funnelform, 10–14 mm, exserted beyond calyx margin; lobes obovate-oblong, apex rounded to truncate or notched. |
Fruiting pedicels | 15–35(–40) mm, sparsely stipitate-glandular to short glandular-villous. |
divergent at right angles from stem, usually closely paired, 10–45 mm, negatively phototropic, causing capsules to be pressed against a cliff face or crevice at time of dehiscence, glandular-puberulent to glandular-villous, hairs 0.1–0.8 mm, vitreous, flattened, multicellular, gland-tipped. |
Fruiting calyces | usually purple-tinged and purple-dotted, broadly campanulate, inflated, sagittally compressed, 11–15 mm, glabrous or sparsely stipitate-glandular to short glandular-villous, villous at sinuses, throat closing, lobes broadly ovate, abaxial usually longer than lateral, adaxial at least 2 times as long as others. |
angled, tubular-campanulate, slightly inflated, 5–7 mm, margins distinctly toothed or lobed, sparsely stipitate-glandular, lobes pronounced, erect. |
Capsules | included, 5–7 mm. |
included, 3–6 mm. |
Anthers | included, glabrous. |
included, glabrous. |
2n | = 28, 56. |
= 32. |
Erythranthe tilingii |
Erythranthe hymenophylla |
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Phenology | Flowering Jul–Sep. | Flowering Apr–Aug(–Sep). |
Habitat | Seeps, springs, stream banks, shallow rivulets, cliff bases, ledges and crevices, steep gravelly slopes, wet meadows. | Steep, seasonally moist, basalt cliffs with west or southwest exposure, mesic coniferous forests. |
Elevation | 1400–3400 m. (4600–11200 ft.) | 800–1300 m. (2600–4300 ft.) |
Distribution |
AZ; CA; ID; MT; NV; OR; UT; AB
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ID; MT; OR |
Discussion | Plants of Erythranthe tilingii are characterized by their relatively low stature and stems arising from a system of thin rhizomes and producing mostly one to three large flowers each; they usually occur at relatively high elevations. Erythranthe tilingii sometimes has been considered to include one or several infraspecific entities; from within this taxonomic amalgam, four distinct species are recognized here: E. caespitosa, E. corallina, E. minor, and E. tilingii. Erythranthe corallina and E. minor probably are more closely related to E. guttata. The populations identified here as E. tilingii from northeastern Oregon northeast to Alberta and southeast to Utah may prove to be a separate (undescribed) species. Erythranthe tilingii in the strict sense is relatively widespread over the western United States and is sympatric with E. caespitosa and E. corallina. Leaves in E. tilingii are variable in size, and particularly in Idaho, they may approach the small size of those of E. caespitosa, but the leaf margins of E. tilingii are distinctly toothed, and the stems are taller and more erect. Across the range of the species, plants sometimes produce very large leaves, but these often occur on plants with characteristically smaller leaves. This wide variability in size apparently does not occur in E. caespitosa. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
In the protologue, R. J. Meinke observed that plants of Erythranthe hymenophylla have reflexed fruiting pedicels that increase seed dispersal back onto the vertical cliff wall, the characteristic habitat of the species. The hanging habit of E. hymenophylla is reflected in a sharp (90º to 180º) bend in the basal nodes and the long pedicels that are closely paired and divergent in parallel at about right angles from the stem. The species also is characterized by it very short calyx to corolla length, relatively short capsules, and large seeds. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 408. | FNA vol. 17, p. 397. |
Parent taxa | Phrymaceae > Erythranthe | Phrymaceae > Erythranthe |
Sibling taxa | ||
Synonyms | Mimulus tilingii, M. caespitosus var. implexus, M. implexus, M. implicatus, M. langsdorffii var. tilingii, M. lucens, M. veronicifolius | Mimulus hymenophyllus |
Name authority | (Regel) G. L. Nesom: Phytoneuron 2012-39: 44. (2012) | (Meinke) G. L. Nesom: Phytoneuron 2012-39: 38. (2012) |
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