Erythranthe tilingii |
Erythranthe hardhamiae |
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larger mountain monkeyflower, mountain monkey-flower, Tiling's monkey-flower |
Hardham's monkeyflower, Santa Lucia monkeyflower |
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Habit | Perennials, rhizomatous, solitary to weakly colonial, rhizomes forming a mass, yellowish, branching, filiform. | Annuals, taprooted. |
Stems | erect-ascending, usually freely branched, 2–35 cm, glabrous or sparsely stipitate-glandular to short glandular-villous. |
erect, simple or branched from basal nodes, (2–)3–13 cm, glabrous or minutely puberulent. |
Leaves | cauline; petiole 0–25 mm, distals 0 mm; blade palmately 3–5-veined, ovate to lanceolate-triangular or narrowly lanceolate (broadly ovate in large-leaved forms), 5–35(–55) mm, base cuneate to attenuate, margins irregularly denticulate, apex acute to obtuse or rounded, surfaces glabrous, sparsely stipitate-glandular to short glandular-villous, glabrate, or sparsely to moderately villous, hairs thick-vitreous, eglandular. |
cauline, basal not persistent; petiole 0 mm; blade palmately 3-veined (in broader ones), linear to oblanceolate, 2–12 × 1–5 mm, base attenuate, margins entire, sometimes toothed, apex acute, surfaces glabrous or minutely puberulent. |
Flowers | herkogamous, 1–3(–5), from distal nodes. |
herkogamous, 1–12, from distal or medial to distal nodes. |
Styles | hirtellous. |
glabrous. |
Corollas | yellow, red-dotted, bilaterally symmetric, bilabiate; tube-throat broadly funnelform, 15–28 mm, exserted 5–10 mm beyond calyx margin; limb expanded 14–30 mm. |
pink to purple, abaxial limb with 2 yellow palate ridges, bilaterally symmetric, strongly bilabiate; tube-throat funnelform, 5–10 mm, exserted 2–5 mm beyond calyx margin; limb expanded 7–13 mm, lobes notched, abaxial limb sparsely bearded. |
Fruiting pedicels | 15–35(–40) mm, sparsely stipitate-glandular to short glandular-villous. |
ascending to often spreading horizontally, 10–60 mm. |
Fruiting calyces | usually purple-tinged and purple-dotted, broadly campanulate, inflated, sagittally compressed, 11–15 mm, glabrous or sparsely stipitate-glandular to short glandular-villous, villous at sinuses, throat closing, lobes broadly ovate, abaxial usually longer than lateral, adaxial at least 2 times as long as others. |
becoming reddish, sometimes red-spotted, campanulate to cylindric, 4–6 mm, margins distinctly toothed or lobed, glabrous or minutely puberulent, ribs weak, lobes pronounced, erect, margins glabrous. |
Capsules | included, 5–7 mm. |
included, 4–5 mm. |
Anthers | included, glabrous. |
included or slightly exserted, glabrous. |
Stigmas | equal in length to corolla tube or exserted. |
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2n | = 28, 56. |
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Erythranthe tilingii |
Erythranthe hardhamiae |
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Phenology | Flowering Jul–Sep. | Flowering Mar–Apr. |
Habitat | Seeps, springs, stream banks, shallow rivulets, cliff bases, ledges and crevices, steep gravelly slopes, wet meadows. | Sandy soils near sandstone outcrops and chaparral. |
Elevation | 1400–3400 m. (4600–11200 ft.) | 300–800 m. (1000–2600 ft.) |
Distribution |
AZ; CA; ID; MT; NV; OR; UT; AB
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CA |
Discussion | Plants of Erythranthe tilingii are characterized by their relatively low stature and stems arising from a system of thin rhizomes and producing mostly one to three large flowers each; they usually occur at relatively high elevations. Erythranthe tilingii sometimes has been considered to include one or several infraspecific entities; from within this taxonomic amalgam, four distinct species are recognized here: E. caespitosa, E. corallina, E. minor, and E. tilingii. Erythranthe corallina and E. minor probably are more closely related to E. guttata. The populations identified here as E. tilingii from northeastern Oregon northeast to Alberta and southeast to Utah may prove to be a separate (undescribed) species. Erythranthe tilingii in the strict sense is relatively widespread over the western United States and is sympatric with E. caespitosa and E. corallina. Leaves in E. tilingii are variable in size, and particularly in Idaho, they may approach the small size of those of E. caespitosa, but the leaf margins of E. tilingii are distinctly toothed, and the stems are taller and more erect. Across the range of the species, plants sometimes produce very large leaves, but these often occur on plants with characteristically smaller leaves. This wide variability in size apparently does not occur in E. caespitosa. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Erythranthe hardhamiae is endemic to the central coast region in Monterey and San Luis Obispo counties. The species was previously included in E. palmeri but can be distinguished by having a wider limb (16–25 mm) than E. palmeri (8–15 mm) and pale pink flowers with a broad yellow palate and orifice. In contrast, E. palmeri has deep pink flowers with two yellow ridges on the palate. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 408. | FNA vol. 17, p. 388. |
Parent taxa | Phrymaceae > Erythranthe | Phrymaceae > Erythranthe |
Sibling taxa | ||
Synonyms | Mimulus tilingii, M. caespitosus var. implexus, M. implexus, M. implicatus, M. langsdorffii var. tilingii, M. lucens, M. veronicifolius | |
Name authority | (Regel) G. L. Nesom: Phytoneuron 2012-39: 44. (2012) | N. S. Fraga: Aliso 30: 64, figs. 23–25. (2012) |
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