Erythranthe tilingii |
Erythranthe grayi |
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larger mountain monkeyflower, mountain monkey-flower, Tiling's monkey-flower |
Gray's monkeyflower, mariposa monkeyflower |
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Habit | Perennials, rhizomatous, solitary to weakly colonial, rhizomes forming a mass, yellowish, branching, filiform. | Annuals, fibrous-rooted. |
Stems | erect-ascending, usually freely branched, 2–35 cm, glabrous or sparsely stipitate-glandular to short glandular-villous. |
erect, simple or branched from base, weakly 4-angled, 8–20 cm, glabrous. |
Leaves | cauline; petiole 0–25 mm, distals 0 mm; blade palmately 3–5-veined, ovate to lanceolate-triangular or narrowly lanceolate (broadly ovate in large-leaved forms), 5–35(–55) mm, base cuneate to attenuate, margins irregularly denticulate, apex acute to obtuse or rounded, surfaces glabrous, sparsely stipitate-glandular to short glandular-villous, glabrate, or sparsely to moderately villous, hairs thick-vitreous, eglandular. |
basal and cauline; petiole 0 mm; blade palmately 3–5-veined, broadly ovate, 7–18 × 5–12 mm, base rounded, margins denticulate, apex acute, surfaces glabrous, rarely sparsely puberulent. |
Flowers | herkogamous, 1–3(–5), from distal nodes. |
herkogamous, 2–20, from proximal to distal nodes. |
Styles | hirtellous. |
glabrous. |
Corollas | yellow, red-dotted, bilaterally symmetric, bilabiate; tube-throat broadly funnelform, 15–28 mm, exserted 5–10 mm beyond calyx margin; limb expanded 14–30 mm. |
rose red, throat pink lined with rose red and a yellow patch, abaxial ridges yellow, bilaterally symmetric, weakly bilabiate; tube-throat cylindric-funnelform, 8–11 mm, exserted beyond calyx margin; limb expanded 7–10 mm. |
Fruiting pedicels | 15–35(–40) mm, sparsely stipitate-glandular to short glandular-villous. |
6–7 mm, shorter than subtending leaves, glabrous. |
Fruiting calyces | usually purple-tinged and purple-dotted, broadly campanulate, inflated, sagittally compressed, 11–15 mm, glabrous or sparsely stipitate-glandular to short glandular-villous, villous at sinuses, throat closing, lobes broadly ovate, abaxial usually longer than lateral, adaxial at least 2 times as long as others. |
campanulate, 9–11 × 5–6 mm, margins subtruncate, glabrous, sometimes densely papillate at flowering with tiny, 1-celled, eglandular hairs, these apparently deciduous by fruiting, lobes reduced, subequal. |
Capsules | included, 5–7 mm. |
included, 5–9 mm. |
Anthers | included, glabrous. |
included, minutely villous-hirsute. |
2n | = 28, 56. |
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Erythranthe tilingii |
Erythranthe grayi |
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Phenology | Flowering Jul–Sep. | Flowering May–Jul(–Oct). |
Habitat | Seeps, springs, stream banks, shallow rivulets, cliff bases, ledges and crevices, steep gravelly slopes, wet meadows. | Drying pond beds, creek banks, yellow pine, yellow pine-Libocedrus woodlands. |
Elevation | 1400–3400 m. (4600–11200 ft.) | 1000–1900 m. (3300–6200 ft.) |
Distribution |
AZ; CA; ID; MT; NV; OR; UT; AB
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CA
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Discussion | Plants of Erythranthe tilingii are characterized by their relatively low stature and stems arising from a system of thin rhizomes and producing mostly one to three large flowers each; they usually occur at relatively high elevations. Erythranthe tilingii sometimes has been considered to include one or several infraspecific entities; from within this taxonomic amalgam, four distinct species are recognized here: E. caespitosa, E. corallina, E. minor, and E. tilingii. Erythranthe corallina and E. minor probably are more closely related to E. guttata. The populations identified here as E. tilingii from northeastern Oregon northeast to Alberta and southeast to Utah may prove to be a separate (undescribed) species. Erythranthe tilingii in the strict sense is relatively widespread over the western United States and is sympatric with E. caespitosa and E. corallina. Leaves in E. tilingii are variable in size, and particularly in Idaho, they may approach the small size of those of E. caespitosa, but the leaf margins of E. tilingii are distinctly toothed, and the stems are taller and more erect. Across the range of the species, plants sometimes produce very large leaves, but these often occur on plants with characteristically smaller leaves. This wide variability in size apparently does not occur in E. caespitosa. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
In addition to the features noted in the key and descriptions, the fruiting calyces of Erythranthe grayi are distinctly more inflated than those of E. acutidens and E. inconspicua. G. L. Nesom (2012g) maintained E. grayi as distinct from E. acutidens, relying primarily on fruiting pedicel length (see key above), but the two have nearly identical ranges (Tuolumne County south to Kern County), and study of additional collections suggests that only a single species may be represented. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 408. | FNA vol. 17, p. 382. |
Parent taxa | Phrymaceae > Erythranthe | Phrymaceae > Erythranthe |
Sibling taxa | ||
Synonyms | Mimulus tilingii, M. caespitosus var. implexus, M. implexus, M. implicatus, M. langsdorffii var. tilingii, M. lucens, M. veronicifolius | Mimulus grayi |
Name authority | (Regel) G. L. Nesom: Phytoneuron 2012-39: 44. (2012) | (A. L. Grant) G. L. Nesom: Phytoneuron 2012-39: 34. (2012) |
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